Authors

  • Farrukh Matkarimov
    PhD student, Institute of Genetics and Plant Experimental Biology, Academy of Sciences of Uzbekistan
  • Oybek Kholliyev
    PhD student, Institute of Genetics and Plant Experimental Biology, Academy of Sciences of Uzbekistan
  • Abdulla Fayzullaev
    Senior Researcher, Institute of Genetics and Plant Experimental Biology, Academy of Sciences of Uzbekistan
  • Oygul Raulova
    Teacher, Chirchik State Pedagogical University, Uzbekistan
  • Dilafruz Kulmamatova
    Head of laboratory, Institute of Genetics and Plant Experimental Biology, Academy of Sciences of Uzbekistan
  • Saidmurat Baboev
    Professor, Institute of Genetics and Plant Experimental Biology, Academy of Sciences of Uzbekistan

DOI:

https://doi.org/10.37547/tajabe/Volume06Issue10-06

Keywords:

Chickpeas microbial fertilizer chlorophyll

Abstract

A field experiment was conducted to study the effects of Rhizobium 3 , Rhizobium 9 and PlantaStim microbial fertilizers on the morphological, physiological and valuable economic characters of chickpea (Сicer arietinum L.). All samples treated with microbial fertilizers showed positive results compared to the control variation. Efficiency of Rhizobium 3, among the microbial fertilizers, was higher. In the chickpea samples, treatment with Rhizobium 3 increased the total chlorophyll content by 14.91%, carotenoid content by 20%, plant biomass by 31.39%, grain number by 19.37%, productivity by 27.25%, total protein content by 25.29%, and amount of nodule by 2.07 times, compared to the control variation.


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PUBLISHED DATE: - 30-10-2024
DOI: -

https://doi.org/10.37547/tajabe/Volume06Issue10-06

PAGE NO.: - 32-42

INFLUENCE OF MICROBIAL FERTILIZERS ON
MORPHOLOGICAL, PHYSIOLOGICAL AND
VALUABLE ECONOMIC CHARACTER OF
CHICKPEA (CICER ARIETINUM L.)


Farrukh Matkarimov

PhD student, Institute of Genetics and Plant Experimental Biology, Academy of Sciences of

Uzbekistan

Oybek Kholliyev

PhD student, Institute of Genetics and Plant Experimental Biology, Academy of Sciences of

Uzbekistan

Abdulla Fayzullaev

Senior Researcher, Institute of Genetics and Plant Experimental Biology, Academy of Sciences

of Uzbekistan

Oygul Raulova

Teacher, Chirchik State Pedagogical University, Uzbekistan

Dilafruz Kulmamatova

Head of laboratory, Institute of Genetics and Plant Experimental Biology, Academy of Sciences

of Uzbekistan

Saidmurat Baboev

Professor, Institute of Genetics and Plant Experimental Biology, Academy of Sciences of

Uzbekistan

RESEARCH ARTICLE

Open Access


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INTRODUCTION

Global climate change is causing water scarcity in

many territories of the world. Ensuring the food

security of the population requires the wide
implementation of high caloric crop types which

are resistant to drought. Chickpea plant is

considered as one of such crops (Field CB, 2014).
The annual production of chickpeas worldwide

reaches up to 15.0 million tons. Herein India’s

makes up 73% (FAOSTAT). There are mainly two

types of chickpeas in the market. They are Kabuli
and Desi.
Desi is cultivated in 75% of the whole chickpea

cultivation area in the world, and 25% of the area
belongs to Kabuli ecotype (Kassie M., 2009).

Chickpea plant has types with large, small and

medium sized seeds, with different colors.
According to the size of seeds, they are divided into

2 groups: large-grained (12 x 9 mm) and small-
grained types. The kabuli type of chickpeas has

large white grains, while the grains of desi type
plant are small, dark in colour (A.Frimpong, 2009).
Chickpea plant grain occupies an important place

in the human diet. It contains protein,

carbohydrate,

folate,

β

-carotene, minerals,

vitamins and fatty acids necessary for human

health (A.K.Jukanti, 2012).
A number of microbial fertilizers are used in the

cultivation of legumes. One of these are microbial

fertilizers based on Rhizobium strains.
Rhizobium bacteria is effective in increasing the

yield of leguminous plants and soil fertility . It is

advisable to use bacterial fertilizers with the most

active strains of the nodule bacteria on leguminous
(mung bean, soybean, bean, lentil, pea) crops

(H.H.Zahran, 1991).
When microbial fertilizers are used in the

cultivation of leguminous crops, the microflora that
mobilizes nitrogen, phosphorus and potassium in

the soil is activated. As a result, the plant will be
able to get the nutritional minerals it needs, not

only during the application period, but also during
the entire vegetation period (D.M.Sytnikov 2012).
Growth and development of leguminous plants is

improved and productivity increases when they
are treated with plant growth-accelerating

rhizobacteria compared to the treatment of

leguminous plants with Rhizobium bacteria
separately (M.S.Dardanelli, 2008; M.Yadegari

2010). Microbial fertilizers obtained on the basis of
azotobacters increase the germination of seeds and

absorption of nutrients in plants . Moreover, it
accelerates the synthesis of protein and amino

acids, benefits the growth of crops and helps
restore soil fertility (A.D.Jnawali, 2015).
Application of Rhizobium and Azotobacter strains

together on chickpea plants increased the amount

of nitrogen in the roots and grain yield as well (A.
Abdiev, 2019; A. Siddiq, 2014).
In addition, there are microbial fertilizers obtained

on the basid of Trichoderma fungi types.
Trichoderma species are symbiotically associated

with the apoplastic part of plant roots and directly
affect the plant. It has a positive effect on the

Abstract


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germination of seeds, absorption of substances and
increases productivity (B.N.Singh 2015).
Microbial fertilizers affect physiological and

biochemical processes in plants, improve plant

growth and development, and serve to increase
crop productivity. Based on this, the aim of the

research is to study the effect of microbial
fertilizers

on

the

change

of

the

morphophysiological and valuable economic
characteristics of the chickpea plant.

MATERIALS AND METHODS

Malkhotra variety of Chickpea (Cicer arietinum L.)

seed was used in the field experiments. Rhizobium

3 and Rhizobium 9 preparations were obtained
from the collection of the Department of

Microbiology

and

Biotechnology,

National

University of Uzbekistan. PlantaStim (Trichoderma

Lignorum) was obtained from private company of
AnGuzal Agroservice, Uzbekistan.
Field experiments were conducted in 2021 and

2022 to study the effect of Rhizobium 3, Rhizobium

9 and PlantaStim on growth, nodulation and yield
of Chickpea (Cicer arietinum L.) at the Institute of

Genetics and Plant Experimental Biology, Kibray,
Tashkent region, Uzbekistan. In the experiments

randomized block design was used with three
replications, in a plot of 10 m2 with a row spacing

of 30 cm and a plant spacing of 10 cm was used.
Experimental treatments included a non-

inoculated control, inoculated with Rhizobium 3,
Rhizobium 9 and PlantaStim.
The soil of the area where the field experiments

were conducted is considered to be irrigated gray

soil, the humus content is 0.8-1.2%, and the level of

availability of mobile phosphorus is on average 30-
38 mg/kg (D.E.Qulmamatova, 2022).
In the Malkhotra variety of chickpea, physiological

indicators were determined during the period of

general flowering of plants.
Extraction and determination of pigment

concentration: Amount of pigments in leaves of

chickpea plant samples were determined in the
experiment, i.e. 3-4 leaves, calculating from the

glineth point. Fifty milligrams of each leaf sample,
placed in a test tube, obtained homogenization in 5

ml of 95% ethyl alcohol solution (Lichtenthaler and
a Wellburn, a 1983). The homogenization

underwent centrifugation at a speed of 5000 rpm

for 12 min. The amount of chlorophyll a, b and
carotenoids in the resulting extract attained

determination by an Agilent Cary 60 UV-Vis
spectrophotometer at 664, 649 and 470 nm. Based

on these indicators, the following equations were
used in calculating the amounts of chlorophyll a, b

and carotenoids in chickpea leaves (Nayek et al.,
2014):
Ch-a=13.36A664

5.19 A649

Ch-b=27.43A649

8.12 A664

C x+c=(1000A470

2.13Ca

97.63Cb)/209

Transpiration rate was determined according to

Ivanov’s method [Ivanov A.A. et.al, 1950], water

holding capacity was determined with the method

by Kushnirenko (Kushnirenko M.D. et al, 1970),

and Tretyakov’s method (

Tretyakov N.N. et.al,

1990) was used in determining the total amount of
water in leaves.
Harvest index was calculated according to the

following formula:

𝐻𝑎𝑟𝑣𝑒𝑠𝑡 𝑖𝑛𝑑𝑒𝑥(%) =

𝑆𝑒𝑒𝑑 𝑦𝑖𝑒𝑙𝑑

𝐵𝑖𝑜𝑙𝑜𝑔𝑖𝑐𝑎𝑙 𝑦𝑖𝑒𝑙𝑑

× 100

Determination of nitrogen and total protein

contents. Kjeldahl is one of the methods used to

determine the amount of total proteins. According
to this method, the total calculated protein has the

amount of nitrogen determining it. The essence of

the process is to hydrolyze the organic substances
in the sample with the help of concentrated sulfuric

acid (amine groups in the protein) to form
ammonium sulfate salts.

nitrogenous organic matter + H

2

SO

4

(NH

4

)

2

SO

4

+ CO

2

+ H

2

O

After completing the hydrolysis, the ammonium sulfate formed receives treatment with sodium


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hydroxide to convert it to ammonia.

2H

2

O

(NH

4

)

2

SO

4

+ 2NaOH Na

2

SO

4

+ 2NH

4

OH

NH

3

Ammonia or ammonium hydroxide formed, after

neutralization, assimilates into the sulfuric acid

solution. The remaining acid gets titrated with an
alkaline solution. Estimating the amount of

nitrogen from the amount of ammonia calculated
followed. An accurate sample for analysis came

from the average crushed homogeneous specimen
of the studied material in a test tube; the error rate

should not exceed 0.1%. The sampling progressed
to quantitate in a Kjeldahl flask. Later, the

experiment advanced according to the protocol
and instructions of Metody Kontrolya (Chemical

Factors, 2004).
Processing of the obtained results. The mass

fraction of nitrogen (X) in the analyzed sample for
calculation used the formula as a percentage of the

mass of the specimen by the volume after the
titration of the amount of ammonia that has passed

through the diluted sulfuric acid.

The volume of 0,1 mol/l sodium hydroxide solution

served to titrate the remaining 0.1 mol/l sulfuric
acid solution in sample experiment vo, ml.
The obtained data were analyzed on the statistical

software

ANOVA

STATGRAPHICS-18

(www.statgraphics.com).
The research was carried out in order to study the

effect of microbial fertilizers on changes in

photosynthetic

pigments,

water

balance,

morphological characters, productivity elements in

chickpea plants.

RESULTS AND DISCUSSIONS

When photosynthetic pigments in leaves were

analyzed, the chickpea plants, which were
inoculated with microbial fertilizers, showed

higher results than the control variation (Figure 1).
In this case, the amount of chlorophyll a increased

by 7.29 (10.94%), the amount of chlorophyll b by
12.05 (24.10%), the total amount of chlorophyll by

7.72 (14.91%) and the amount of carotenoids by
15, 55 (20%) as well. No difference was observed

between the amounts of photosynthetic pigments
in samples treated with Rhizobium 9 and

PlantaStim microbial fertilizers. The samples,

which were treated with Rhizobium 3, chlorophyll
a, chlorophyll b, total chlorophyll, and carotenoid,

showed the highest indicators.


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Figure 1. The effect of microbial fertilizers on changes in the amount of photosynthetic pigments

in chickpea plants

Changes in the rate of photosynthesis are directly

related to the photosynthetic pigments in
chloroplasts (Maisura, et al., 2014). These pigments

affect the growth and development of the plant
(L.Taiz, et al., 2006). A number of abiotic factors

influence the change in the amount of

photosynthetic pigments. The decrease in the
amount of chlorophyll in the leaves of plants is

mainly observed in conditions of salinity stress
(Z.A.Saqib, 2012), heat stress (S.Sangwan, 2012),

water stress (soil waterlogging) (Amri M., 2014)
and water shortage (Y.Alaei, 2011).
Mineral fertilizers, especially nitrogen fertilizers,

play an important role in increasing the amount of
chlorophyll in plant leaves. A high chlorophyll

content results in a high yield (I.Skudra, 2017).

Moreover, a number of microbial fertilizers also
cause changes in the amount of photosynthetic

pigments in leaves. In soybean, Rhizotorphin
microbial fertilizer (Yu.Tsvetkova, 2020), in green

pea (Pisum sativum L.), microbial fertilizer
obtained from T42 strain of Trichoderma

asperellum (B.N.Singh, 2015) and in mung bean

Rhizobium 3 and Rhizobium 9 (Matkarimov, 2024)
increased chlorophyll and carotenoids content up

to 10.91%-14.08% respectively.
Influence of microbial fertilizers on the

physiological processes as transpiration rate, total

amount of water in leaves, water retention

property, which are related to water exchange in
chickpea plant, was comparatively analyzed (Table

1). According to the results of the analysis, the
transpiration rate in the leaves of chickpea plants,

treated with Rhizobium 3 and Rhizobium 9, did not
differ significantly from the control variations. It

helped to partially increase the total amount of
water in leaves and water retention property.
In the samples, treated with PlantaStim, the

transpiration rate increased by 9.95%, and the

total amount of water in leaves and water retention
property decreased partially. In the process of

transpiration, the increase of absorption of water
and mineral substances, with the help of root hairs,

and evaporation from the leaves, lead to decrease
in temperature and prevent overheating of the

leaves (Sagdiev M.T., 2007).

Table 1

The effect of microbial fertilizers on physiological processes related to water exchange in

chickpea plants

1.92

2.13

2.04

2.06

0.83

1.03

0.92

0.93

2.75

3.16

2.96

2.98

0.45

0.54

0.52

0.52

0

0.5

1

1.5

2

2.5

3

3.5

Сontrol

Rhizobium 3

Rhizobium 9

PlantaStim

A

m

o

u

n

t

o

f

p

h

o

to

sy

n

th

eti

c

p

ig

m

en

ts,

m

g

/g

Chlorophyll "a"

Chlorophyll "b"

Total chlorophyll

Carotenoids


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Transpiration rate

(mg Н

2

O /1 g. wet

leaf x 1 hour)

Total amount

of water in

leaves,%

Water retention

property,%

Monitor

277.33 ± 5.48

80.87 ± 0.33

31.09 ± 0.28

Rhizobium 3

279.75 ± 7.16

81.37 ± 0.28

30.16 ± 0.18

Rhizobium 9

280.26 ± 5.25

80.99 ± 0.31

30.23 ± 0.22

PlantaStim

304.93 ± 8.16

79.21 ± 0.20

32.37 ± 0.52

Height of one plant 83.47±0.91 cm in control variations, and under variations treated with Rhizobium 3,

Rhizobium 9 and PlantaStim was 86.47±0.57 cm, 85.33±0.88 and 84.07 cm respectively (Fig. 2). There
wa

sn’t noted significant difference in plant height among the variations. However, the highest value was

detected under the variation treated with Rhizobium 3, and it was 3.59% higher than control variation.

Figure 2. Effect of microbial fertilizers on plant height of pea plants

Plant height and branching are important morphological characteristics of chickpea plant (N.S.Taspaev,

2018). Primary and secondary branching is considered to be one of the main morphological characters
in chickpea plant. It was observed that the number of main branches of one plant was from 3.20±0.14 to

3.33±0.13, and the number of side branches was from 14.87±0.40 to 15.47±0.31. The highest rate was
found under the variation treated with Rhizobium 3 (Fig. 3).

83.47

86.47

85.33

84.07

80

81

82

83

84

85

86

87

88

Control

Rhizobium 3

Rhizobium 9

PlantaStim

P

lan

t h

eig

h

t,

cm

3.2

3.33

3.27

3.33

14.87

15.47

15.27

15.27

0

5

10

15

20

Control

Rhizobium 3

Rhizobium 9

PlantaStim

Primary branches per plant

Secondary branches per plant


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Figure 3. Effect of microbial fertilizers on branching of chickpea plants

Chickpea is a plant with a deep and strong root

system, which forms nodules in the roots together

with Rhizobium bacteria. These nodules have the

property of fixing atmospheric nitrogen in a useful
form for the plant. In chickpea, nodulation begins

approximately one month after germination
(P.M.Gaur, 2010). The number of nodules was

higher in chickpea plant samples treated with
microbial fertilizers as Rhizobium 3 and Rhizobium

9 compared to control variation and PlantaStim
microbial fertilizer (Fig. 4). In this case, the nodule

mass per plant during the general flowering phase
was 2.22±0.05 g. and 2.21±0.05 g. in the control

variation and plants treated with PlantaStim,

respectively, and in the samples treated with

Rhizobium 3 and Rhizobium 9 the nodule mass was
4.60±0.17 g and 3.84±0.09 g, respectively. The root

nodule mass in the treated samples with
Rhizobium 3 and Rhizobium 9 was 2.07 and 1.73

times higher than in the control variety,
respectively (Fig. 5). An increase in nodule mass in

the root of a chickpea plant leads to an increase in

nitrogen fixation (Wadisirisuk, 1985). It has been
found that treating mung bean plants with

Rhizobium 3 and Rhizobium 9 microbial fertilizers
has a positive effect on the formation of root

nodules, growth and development (F.Matkarimov
2019).


Rhizobium 9

Rhizobium 3

PlantaStim

Control

Figure 4. Formation of nodules in chickpea plants as a result of the effect of microbial fertilizers.


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Figure 5. Effect of microbial fertilizers on changes in nodule mass in chickpea plants

Increasing crop productivity in agriculture is of great importance. In the framework of the research,

valuable economic characteristics such as biomass per plant, number of pods per plant, number of grains

per plant, pods weight per plant, grain weight per plant, 100-seed weight, grain yield (1 m2/g), harvest
index and total protein quantity were analyzed (Table 2).

Table 2

Effect of microbial fertilizers on valuable economic characteristics of chickpea plant

Control

Rhizobium 3

Rhizobium 9

PlantaStim

Biomass per plant, g.

53.74±1.79

70.61±1.65

63.68±1.19

58.57±1.57

Number of

pods

per plant

68.20±2.05

76.80±1.46

73.40±0.86

70.67±1.46

Number

of

grains per plant

75.73±1.80

90.4±2.09

88.20±1.47

84.87±1.3

Pod

weight

per plant, g.

35.93±1.06

40.12±1.02

38.04±0.80

37.36±0.70

Grains

weight

per plant, g.

24.04±0.56

30.31±0.68

28.30±0.66

28.33±0.43

100-seed weight

33.07±0,54

33.68±0,41

33.15±0,48

33,41±0,61

Grain yield, g. (1 m

2

)

315.16±9,1

401.04±8,6

376.48±9,4

377.88±10,7

Total protein content, %

21.43

26.8 5

24.13

23.37

Harvest index

45.08

43.06

44,42

44.60

The above-ground dry biomass indicator of the

chickpea in one plant was 53.74±1.79 g in control
variation, and in the samples treated with

Rhizobium 3, Rhizobium 9 and PlantaStim, this
indicator showed 70.61±1.65 g, 63.68±1.19 g and

58.57±1.57 g, respectively. The main yield

elements as the number of pods per plant, the
number of grains, the weight of pods and the

weight of grains per plant were 68.20±2.05 pcs,
75.73±1.80 pcs, and 35.93±1.06 g, and 24.04±0.56

g, respectively. These indicators were higher in all
variations treated with microbial fertilizers than

the control variation. The grain weight of the plant
is of great importance in obtaining high yield (S.V.

Bulyntsev, 2015). The main components that
improve grain yield in chickpea plants are plant

2.22

4.6

3.84

2.21

0

1

2

3

4

5

6

Control

Rhizobium 3

Rhizobium 9

PlantaStim

No

d

u

le

m

ass

,

g

.


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biomass, grain number and grain weight
(Qulmamatova, D.E. 2023). According to the results

of the experiment the weight of 100 grains made up
33.07 - 33.68 g. Grain weight of the samples treated

with microbial fertilizers increased slightly. The
grain yield per 1 m2 under the control variety was

315.16 g, whereas, in the samples treated with
Rhizobium 3, Rhizobium 9 and PlantaStim was

401.04 g, 376.48 g, and 377.88 g, respectively. The

highest rate was observed in the samples treated
with Rhizobium 3 and the yield was 27.25% higher

than in the control variation. The total amount of
protein in the grain was between 21.43 and

26.85%. Also, the highest rate was observed in the
samples treated with Rhizobium 3, and the yield

was 25.29% higher than in the control variety. The
harvest index was 43.06 - 45.08. Higher biomass in

the samples treated with microbial fertilizers led to
a partial decrease in the yield index.
In the chickpea samples grown under field

conditions, higher total water amount, number of

photosynthetic pigments, root nodules and yield
elements in the leaves treated with Rhizobium 3

and Rhizobium 9 microbial fertilizers, as well as
increased transpiration rate, photosynthetic

pigment content and yield elements in the samples
treated with PlantaStim had a significant positive

effect on the productivity of the plant, compared to
the control variation.
During the selection process, in cultivation of

chickpea, Rhizobium 3 , Rhizobium 9 and

PlantaStim microbial fertilizers have a positive
effect on the morphological, physiological

characteristics of the plant, as well as the valuable
economic characteristics, and serve to increase

crop productivity.

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Patancheru 502 324, Andhra Pradesh, India:

International Crops Research Institute for
the Semi-Arid Tropics. 28 pp.

9.

Ivanov, L. A., Silina, A. A., & Tsel'niker Yu, L.

(1950). O metode bystrogo vzveshivaniya

dlya

opredeleniya

transpiratsii

v

estestvennykh usloviyakh [On the Method of

Rapid Weighing to Determine the
Transpiration

under

the

Natural

Conditions]. Botanicheskiy zhurnal, 35(2),
171-185.

10.

Jnawali, A. D., Ojha, R. B., & Marahatta, S.


background image

THE USA JOURNALS

THE AMERICAN JOURNAL OF AGRICULTURE AND BIOMEDICAL ENGINEERING (ISSN

2689-1018)

VOLUME 06 ISSUE10

41

https://www.theamericanjournals.com/index.php/tajabe

(2015). Role of Azotobacter in soil fertility
and sustainability

a review. Adv. Plants

Agric. Res, 2(6), 1-5.

11.

Jukanti, A. K., Gaur, P. M., Gowda, C. L. L., &

Chibbar, R. N. (2012). Nutritional quality and
health benefits of chickpea (Cicer arietinum

L.): a review. British Journal of Nutrition,
108(S1), S11-S26.

12.

Kassie M., Shiferaw B., Asfaw S. et al. Current

situation and future outlooks of the chickpea
subsector in Ethiopia.

Nairobi: ICRISAT,

2009.

43 p.

13.

Kushnirenko, M. D., Goncharova, E. A., &

Bondar, E. M. (1970). Metody izucheniya
vodnogo obmena i zasukhoustoychivosti

plodovykh rasteniy [Methods for studying
water metabolism and drought tolerance of

fruit plants]. Kishinev: AS MSSR.[in Russian].

14.

Lichtenthaler, H. K., & Wellburn, A. R. (1983).

Determinations of total carotenoids and
chlorophylls a and b of leaf extracts in

different solvents. J. aBiochem.aSoc.aTrans.
11:591-592.

15.

Maisura, et al. "Some physiological character

responses of rice under drought conditions
in a paddy system." J. ISSAAS Vol. 20, No.

1:104-114 (2014)

16.

Matkarimov F., Xakimov A., Rasulova O.,

Tuxtayev D., Qulmamatova D., Baboyev S.

(2024). Mikrobiologik oʻgʻitlarning mosh

(Vigna radiate L.) oʻsimligidagi fot

osintetik

pigmentlar miqdoriga ta’siri. Oziq ovqat

xavfsizligi: Milliy va global muommolar ,(2),
60-64.

17.

Matkarimov, F., Jabborova, D., & Baboev, S.

(2019). Enhancement of plant growth,

nodulation and yield of mungbean (Vigna
radiate L.) by Microbial Preparations.

International

Journal

of

Current

Microbiology and Applied Sciences, 8(08),

2382-2388.

18.

Nayek, S., Choudhury, I.H., Jaishee, N., Roy S.

2014. Spectrophotometric analysis of

chlorophylls

and

carotenoids

from

commonly glinen ferm species by using

various extracting solvents. Res. J. Chem. Sci.
4:63-69.

19.

Qulmamatova D.E., Baboev S.K. and Buronov

A.K. Genetic variability and inheritance

pattern of yield componentsthrough diallel
analysis in spring wheat. SABRAO Journal of

Breeding and Genetics 54 (1) 21-29, 2022
http://doi.org/10.54910/sabrao2022.54.1.

3 http://sabraojournal.org/ pISSN 1029-
7073; eISSN 2224-8978

20.

Qulmamatova, D. E. Chickpea (Cicer

arietinum L.) genotypes evaluation for high
yield through multivariate analysis. SABRAO

Journal of Breeding and Genetics 55 (1) 107-

114,

2023

http://doi.org/10.54910/sabrao2023.55.1.

10

21.

Sangwan, S., Ram, K., Rani, P., & Munjal, R.

(2018). Effect of terminal high temperature

on chlorophyll content and normalized
difference vegetation index in recombinant

inbred lines of bread wheat. Int. J. Curr.
Microbiol. App. Sci, 7(6), 1174-1183.

22.

Saqib, Z. A., Akhtar, J., Ul-Haq, M. A., Ahmad,

I., & Bakhat, H. F. (2012). Rationality of using

various physiological and yield related traits
in determining salt tolerance in wheat.

African Journal of Biotechnology, 11(15),
3558-3568.

23.

Siddiqui, A., Shivle, R., Magodiya, N., &

Tiwari, K. (2014). Mixed effect of Rhizobium

and Azotobacter as biofertilizer on
nodulation and production of chick pea,

Cicer arietinum. Biosci. Biotech. Res. Comm,
7(1), 46-49.

24.

Singh, B. N., Singh, A., Singh, G. S., & Dwivedi,

P. (2015). Potential role of Trichoderma
asperellum T42 strain in growth of pea plant

for sustainable agriculture. J. Pure Appl.

Microbiol, 9(2), 1069-1074.

25.

Skudra, I., & Ruza, A. (2017). Effect of

nitrogen and sulphur fertilization on

chlorophyll content in winter wheat. Rural
sustainability research, 37(332), 29-37.

26.

Taiz, L., & Zeiger, E. (2006). Plant physiology


background image

THE USA JOURNALS

THE AMERICAN JOURNAL OF AGRICULTURE AND BIOMEDICAL ENGINEERING (ISSN

2689-1018)

VOLUME 06 ISSUE10

42

https://www.theamericanjournals.com/index.php/tajabe

4th ed., Sinauer Associates Inc. Publishers,
Massachusetts. 2006. pp.126-128.

27.

Wadisirisuk, P., & Weaver, R. W. (1985).

Importance of bacteroid number in nodules

and effective nodule mass to dinitrogen
fixation by cowpeas. Plant and Soil, 87, 223-

231.

28.

Yadegari, M., & Rahmani, H. A. (2010).

Evaluation of bean(Phaseolus vulgaris)

seeds' inoculation with Rhizobium phaseoli
and

plant

growth

promoting

rhizobacteria(PGPR) on yield and yield
components. African Journal of Agricultural

Research, 5(9), 792-799.

29.

Zahran, H. H. (1991). Conditions for

successful Rhizobium-legume symbiosis in
saline environments. Biology and Fertility of

Soils, 12, 73-80.

30.

Bulynsev, S.V., Novikova, L. Y.U., Gridnev, G.

A.,

&

Sergeyev,

Ye.

A.

(2015).

Korrelyasionnye

svyazi

seleksionnyx

priznakov, opredelyayushix produktivnost
obrazsov nuta (Cicer arietinum L.) iz

kolleksii VIR v usloviyax Tambovskoy
oblasti. Selskoxozyaystvennaya biologiya,

(1), 63-74.

31.

Sagdiyev M.T., Alimova R.A.. Oʻsimliklar

fiziologiyasi. // Oʻquv qoʻllanma. –

Toshkent.

Yangiyoʻl poligraf se

rvis. 2007. 33

bet.

32.

Sytnikov, D. M. (2012). Biotexnologiya

mikroorganizmov

azotfiksatorov

i

perspektivy primeneniya preparatov na ix

osnove. Biotechnologia Achta, 5(4), 034-045.

33.

Taspayev N.S. Produktivnost nuta v

zavisimosti ot srokov poseva, nor

m vыseva i

udobreniy

na

kashtanovыx

pochvax

saratovskogo

zavoljya

//

Dissertatsiya.2018.-str 83

34.

Tretyakov, N. N., Karnauxova, T. V., &

Panichkin, L. A. (1990). Praktikum po
fiziologii rasteniy.

3-ye izd., dop. i pererab.

M.: Agropromizdat,

1990.

271 s.

35.

Svetkova, YU. V., Lyashko, M. U., &

Strajnikova, I. I. (2020). Vliyaniye

primeneniya inokulyanta “Rizotorfin” na

soderjaniye

xlorofilla

v

listyax

introdutsirovannыx sortov soi i ix

urojaynost.

Vestnik

Rossiyskogo

universiteta drujby narodov. Seriya:

Agronomiya i jivotnovodstvo, 15(1), 7-18.

36.

Руководство,

Р.

"4.1.

1672

-03

«Руководство по методам контроля

качества и безопасности биологически
активных добавок к пище»." М.:

Федеральный центр Госсанэпиднадзора
Минздрава России 240 (2004).

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Alaei, Y. (2011). The effect of amino acids on leaf chlorophyll content in bread wheat genotypes under drought stress conditions. Middle-East J. Sci. Res, 10(1), 99-101.

Amri, M., El Ouni, M. H., & Salem, M. B. (2014). Waterlogging affect the development, yield and components, chlorophyll content and chlorophyll fluorescence of six bread wheat genotypes (Triticum aestivum L.). Bulg. J. Agric. Sci, 20(3), 647-657.

Dardanelli, M. S., de Cordoba, F. J. F., Espuny, M. R., Carvajal, M. A. R., Díaz, M. E. S., Serrano, A. M. G., & Megías, M. (2008). Effect of Azospirillum brasilense coinoculated with Rhizobium on Phaseolus vulgaris flavonoids and Nod factor production under salt stress. Soil Biology and Biochemistry, 40(11), 2713-2721.

FAOSTAT. Food and Agriculture Organization of the United Nations. Rome: FAO. 2022.

Field, C. B., & Barros, V. R. (Eds.). (2014). Climate change 2014–Impacts, adaptation and vulnerability: Regional aspects. Cambridge University Press.

Frimpong, A., Sinha, A., Tar'an, B., Warkentin, T. D., Gossen, B. D., & Chibbar, R. N. (2009). Genotype and growing environment influence chickpea (Cicer arietinum L.) seed composition. Journal of the Science of Food and Agriculture, 89(12), 2052-2063.

Gaur PM, Tripathi S, Gowda CLL, Ranga Rao GV, Sharma HC, Pande S and Sharma M. 2010. Chickpea Seed Production Manual. Patancheru 502 324, Andhra Pradesh, India: International Crops Research Institute for the Semi-Arid Tropics. 28 pp.

Ivanov, L. A., Silina, A. A., & Tsel'niker Yu, L. (1950). O metode bystrogo vzveshivaniya dlya opredeleniya transpiratsii v estestvennykh usloviyakh [On the Method of Rapid Weighing to Determine the Transpiration under the Natural Conditions]. Botanicheskiy zhurnal, 35(2), 171-185.

Jnawali, A. D., Ojha, R. B., & Marahatta, S. (2015). Role of Azotobacter in soil fertility and sustainability–a review. Adv. Plants Agric. Res, 2(6), 1-5.

Jukanti, A. K., Gaur, P. M., Gowda, C. L. L., & Chibbar, R. N. (2012). Nutritional quality and health benefits of chickpea (Cicer arietinum L.): a review. British Journal of Nutrition, 108(S1), S11-S26.

Kassie M., Shiferaw B., Asfaw S. et al. Current situation and future outlooks of the chickpea subsector in Ethiopia. – Nairobi: ICRISAT, 2009. – 43 p.

Kushnirenko, M. D., Goncharova, E. A., & Bondar, E. M. (1970). Metody izucheniya vodnogo obmena i zasukhoustoychivosti plodovykh rasteniy [Methods for studying water metabolism and drought tolerance of fruit plants]. Kishinev: AS MSSR.[in Russian].

Lichtenthaler, H. K., & Wellburn, A. R. (1983). Determinations of total carotenoids and chlorophylls a and b of leaf extracts in different solvents. J. aBiochem.aSoc.aTrans. 11:591-592.

Maisura, et al. "Some physiological character responses of rice under drought conditions in a paddy system." J. ISSAAS Vol. 20, No. 1:104-114 (2014)

Matkarimov F., Xakimov A., Rasulova O., Tuxtayev D., Qulmamatova D., Baboyev S. (2024). Mikrobiologik oʻgʻitlarning mosh (Vigna radiate L.) oʻsimligidagi fotosintetik pigmentlar miqdoriga ta’siri. Oziq ovqat xavfsizligi: Milliy va global muommolar ,(2), 60-64.

Matkarimov, F., Jabborova, D., & Baboev, S. (2019). Enhancement of plant growth, nodulation and yield of mungbean (Vigna radiate L.) by Microbial Preparations. International Journal of Current Microbiology and Applied Sciences, 8(08), 2382-2388.

Nayek, S., Choudhury, I.H., Jaishee, N., Roy S. 2014. Spectrophotometric analysis of chlorophylls and carotenoids from commonly glinen ferm species by using various extracting solvents. Res. J. Chem. Sci. 4:63-69.

Qulmamatova D.E., Baboev S.K. and Buronov A.K. Genetic variability and inheritance pattern of yield componentsthrough diallel analysis in spring wheat. SABRAO Journal of Breeding and Genetics 54 (1) 21-29, 2022 http://doi.org/10.54910/sabrao2022.54.1.3 http://sabraojournal.org/ pISSN 1029-7073; eISSN 2224-8978

Qulmamatova, D. E. Chickpea (Cicer arietinum L.) genotypes evaluation for high yield through multivariate analysis. SABRAO Journal of Breeding and Genetics 55 (1) 107-114, 2023 http://doi.org/10.54910/sabrao2023.55.1.10

Sangwan, S., Ram, K., Rani, P., & Munjal, R. (2018). Effect of terminal high temperature on chlorophyll content and normalized difference vegetation index in recombinant inbred lines of bread wheat. Int. J. Curr. Microbiol. App. Sci, 7(6), 1174-1183.

Saqib, Z. A., Akhtar, J., Ul-Haq, M. A., Ahmad, I., & Bakhat, H. F. (2012). Rationality of using various physiological and yield related traits in determining salt tolerance in wheat. African Journal of Biotechnology, 11(15), 3558-3568.

Siddiqui, A., Shivle, R., Magodiya, N., & Tiwari, K. (2014). Mixed effect of Rhizobium and Azotobacter as biofertilizer on nodulation and production of chick pea, Cicer arietinum. Biosci. Biotech. Res. Comm, 7(1), 46-49.

Singh, B. N., Singh, A., Singh, G. S., & Dwivedi, P. (2015). Potential role of Trichoderma asperellum T42 strain in growth of pea plant for sustainable agriculture. J. Pure Appl. Microbiol, 9(2), 1069-1074.

Skudra, I., & Ruza, A. (2017). Effect of nitrogen and sulphur fertilization on chlorophyll content in winter wheat. Rural sustainability research, 37(332), 29-37.

Taiz, L., & Zeiger, E. (2006). Plant physiology 4th ed., Sinauer Associates Inc. Publishers, Massachusetts. 2006. pp.126-128.

Wadisirisuk, P., & Weaver, R. W. (1985). Importance of bacteroid number in nodules and effective nodule mass to dinitrogen fixation by cowpeas. Plant and Soil, 87, 223-231.

Yadegari, M., & Rahmani, H. A. (2010). Evaluation of bean(Phaseolus vulgaris) seeds' inoculation with Rhizobium phaseoli and plant growth promoting rhizobacteria(PGPR) on yield and yield components. African Journal of Agricultural Research, 5(9), 792-799.

Zahran, H. H. (1991). Conditions for successful Rhizobium-legume symbiosis in saline environments. Biology and Fertility of Soils, 12, 73-80.

Bulynsev, S.V., Novikova, L. Y.U., Gridnev, G. A., & Sergeyev, Ye. A. (2015). Korrelyasionnye svyazi seleksionnyx priznakov, opredelyayushix produktivnost obrazsov nuta (Cicer arietinum L.) iz kolleksii VIR v usloviyax Tambovskoy oblasti. Selskoxozyaystvennaya biologiya, (1), 63-74.

Sagdiyev M.T., Alimova R.A.. Oʻsimliklar fiziologiyasi. // Oʻquv qoʻllanma. –Toshkent. Yangiyoʻl poligraf servis. 2007. 33 – bet.

Sytnikov, D. M. (2012). Biotexnologiya mikroorganizmov azotfiksatorov i perspektivy primeneniya preparatov na ix osnove. Biotechnologia Achta, 5(4), 034-045.

Taspayev N.S. Produktivnost nuta v zavisimosti ot srokov poseva, norm vыseva i udobreniy na kashtanovыx pochvax saratovskogo zavoljya // Dissertatsiya.2018.-str 83

Tretyakov, N. N., Karnauxova, T. V., & Panichkin, L. A. (1990). Praktikum po fiziologii rasteniy.–3-ye izd., dop. i pererab. M.: Agropromizdat,–1990.–271 s.

Svetkova, YU. V., Lyashko, M. U., & Strajnikova, I. I. (2020). Vliyaniye primeneniya inokulyanta “Rizotorfin” na soderjaniye xlorofilla v listyax introdutsirovannыx sortov soi i ix urojaynost. Vestnik Rossiyskogo universiteta drujby narodov. Seriya: Agronomiya i jivotnovodstvo, 15(1), 7-18.

Руководство, Р. "4.1. 1672-03 «Руководство по методам контроля качества и безопасности биологически активных добавок к пище»." М.: Федеральный центр Госсанэпиднадзора Минздрава России 240 (2004).