Authors

  • Khamidova Farida Muinovna
    Samarkand State Medical University, Uzbekistan
  • Nurullayev Javohir Asqar O’gli
    Samarkand State Medical University, Uzbekistan

DOI:

https://doi.org/10.37547/ajbspi/Volume04Issue06-09

Keywords:

Morphology skin keratoderma

Abstract

Studying the morphological and genetic different types of keratoses is key for correct diagnosis and choice of treatment. The association between metabolic disorders and skin diseases highlights the need for an integrated approach to the management of patients with chronic diseases, including diabetes mellitus. Further research is needed to better understand the pathogenesis and develop effective treatments.


background image

Volume 04 Issue 06-2024

54


American Journal Of Biomedical Science & Pharmaceutical Innovation
(ISSN

2771-2753)

VOLUME

04

ISSUE

06

P

AGES

:

54-69

OCLC

1121105677
















































Publisher:

Oscar Publishing Services

Servi

ABSTRACT

Studying the morphological and genetic different types of keratoses is key for correct diagnosis and choice of
treatment. The association between metabolic disorders and skin diseases highlights the need for an integrated
approach to the management of patients with chronic diseases, including diabetes mellitus. Further research is
needed to better understand the pathogenesis and develop effective treatments.

KEYWORDS

Morphology, skin, keratoderma, diabetes mellitus.

INTRODUCTION

In our country, data on the epidemiology of

keratoderma is extremely scarce, which is due to a

number of reasons. Firstly, the analysis of statistical

data and publications in the scientific literature is

complicated by the presence of many synonyms for the

disease: seborrheic wart, seborrheic keratoma, senile

wart, basal cell papilloma, pigmented epithelioma,

pigmented

papilloma,

seborrheic

acanthoma,

pigmented basal cell epithelioma, senile papilloma, etc.

Secondly, despite the fact that this benign tumor was

first mentioned by S. Pollitzer at the end of the 19th

century (1890) (1,4,7), its detailed description by D.I.

Golovin did it relatively recently - in 1958 (2,3). And

thirdly, to date the etiology of this proliferative process

Research Article

MORPHOGENETIC

CHARACTERISTICS

OF

KERATODERMA

IN

METABOLIC DISORDERS

Submission Date:

June 20, 2024,

Accepted Date:

June 25, 2024,

Published Date:

June 30, 2024

Crossref doi:

https://doi.org/10.37547/ajbspi/Volume04Issue06-09


Khamidova Farida Muinovna

Samarkand State Medical University, Uzbekistan

Nurullayev Javohir Asqar O’gli

Samarkand State Medical University, Uzbekistan

Journal

Website:

https://theusajournals.
com/index.php/ajbspi

Copyright:

Original

content from this work
may be used under the
terms of the creative
commons

attributes

4.0 licence.


background image

Volume 04 Issue 06-2024

55


American Journal Of Biomedical Science & Pharmaceutical Innovation
(ISSN

2771-2753)

VOLUME

04

ISSUE

06

P

AGES

:

54-69

OCLC

1121105677
















































Publisher:

Oscar Publishing Services

Servi

on the skin is not clear. According to the international

histological class

ification of skin tumors “Pathology

and Genetics of Skin Tumors”, published under the

auspices of the International Agency for Research on

Cancer (5,6,7), seborrheic keratosis is a benign

epithelial tumor and is a type of acanthoma. The

disease affects men and women almost equally,

predominantly over 40 years of age (17). A clear

correlation of its debut with age has been established.

Thus, in the cohort of 24

49 year olds, the prevalence

of seborrheic keratosis was 38%, in 50

59 year olds

69%, in 60

69 year olds

86%, and among 70

79 year

olds

more than 90% (34). The disease is extremely

common in countries with high levels of insolation, for

example, in Australia, where it occurs in 100% of cases

in the population over 50 years of age. (35).

European studies indicate a slightly lower incidence of

this epithelial tumor: 82% among men and 62% among

women aged over 70 years (8,9,10,37,39). Russian

researchers point out that closed areas of the skin

exposed to mechanical stress (friction, pressure) are

most affected. In patients with numerous elements of

seborrheic keratosis, a positive family history is often

recorded (11,12,13,42,49). Seborrheic keratosis is

represented by multiple foci of proliferative growth

(14,16,21,45).

Dermatoscopic

and

histological

examination is mandatory to verify the diagnosis

(15,16,23,28,29).

Seborrheic keratosis (SK) is a common benign tumor of

the epidermis in men and women, occurring more

often after 45-50 years. KS elements appear as distinct

brown spots and/or plaques widely distributed along

the edges of the skin (17,18,19,25,26). There is no

unified morphological classification of this disease,

since the histological manifestations of KS are varied -

symptoms of several histological types may be present

in the same lesion (20,25,28,29).

In foreign literature they are divided into acanthous,

hyperkeratotic, adenoid, irritated, clonal type and

melanoacanthomas. The diagnosis of KS in most cases

is not in doubt, but the tumor can mimic other skin

neoplasms: common warts, lentigines, melanocytic

nevi, actinic keratosis, Bowen's disease, squamous cell

carcinoma both clinically and during pathological

examination (21,22,32,38). The etiology of KS is

unknown, although genetics, increased sun exposure,

somatic mutations in fibroblast growth factor receptor

3 (FGFR3), and human papillomavirus have been

identified as risk factors (23,24,31,35). Currently, the

theory of keratinocyte aging and impaired apoptosis in

KS is widely accepted. Studies are rare, and the

reasons for changes in the proliferative properties of

cells in KS are unclear (25,26,27). In addition, MK is of

interest for studying cell cycle disorders, since

according to modern histological criteria it is classified

as a benign tumor (28,29,30). However, many

immunohistochemical

features

of

malignant

neoplasms

and

clinical

cases

of

malignant

transformation within MC foci make it necessary to

continue the targeted study of the pathogenesis of the


background image

Volume 04 Issue 06-2024

56


American Journal Of Biomedical Science & Pharmaceutical Innovation
(ISSN

2771-2753)

VOLUME

04

ISSUE

06

P

AGES

:

54-69

OCLC

1121105677
















































Publisher:

Oscar Publishing Services

Servi

neoplasm. The main mechanism of proliferation

impairment is a defect in the function of suppressor

genes. In addition to high proliferation, tumor growth

is associated with impaired cell ability to regulate

apoptosis.

The most studied cell cycle markers that are involved in

the control of the proliferative potential of any cell are

p53, p21, p27 and p16. Determination of their

expression level using immunohistochemical methods

allows us to identify the degree of proliferation

disorder, which is invariably accompanied by tumor

growth, the ability of the tumor to invade and

metastasize (10,12,13,14). p27 (Kip1) is an inhibitor of

cyclin-dependent kinase 1B, a product of the human

CDKN1B suppressor gene, a member of the Cip/Kip

protein family. This protein regulates the course of the

cell cycle, is responsible for its arrest in the G1 phase by

suppressing the activity of the cyclin A/cyclin-

dependent kinase 2 and cyclin E/cyclin-dependent

kinase 2 complexes. p27 is a nuclear-cytoplasmic

protein, its intracellular localization is regulated post-

translational modifications. It performs its inhibitory

functions in the nucleus; after its movement into the

cytoplasm, further advancement of the cell through

the cycle becomes possible (10,11,15). Overexpression

of p27 has been studied in malignant neoplasms of

internal organs and is an unfavorable prognostic factor

for tumor progression (24,27,28). There are few studies

on benign neoplasms, and they contain conflicting

data. Thus, in a study of 10 irritated and acanthotic SCs,

A. Brueks et al. (8) overexpression of only p27 was

detected along with the absence of expression of p53,

p16 and a low proliferative index, which allowed the

authors to talk about the leading role of p27 in the

control of cell proliferation. However, it seems to us

appropriate to study p27 in all types of KS, taking into

account their morphological diversity. As a result of the

study, a violation of p27 protein expression was

revealed in all histological types of KS, which indicates

its significant role in the pathogenesis of the disease.

Increased expression of p27, normally found in the

greatest quantity in cells in the G1 phase (19,20), was

found in the authors’ studies in irritable, adenoid and

some cases of clonal types. Considering that p27 is a

“reserve brake” of cell division (12,13), its significant

increase in KS may indicate the absence or insufficient

response of the first link of cell cycle regulators - p53

and p16 to an increase in the proliferative activity of

cells. The staining of nuclear membranes and adjacent

areas of the cytoplasm was considered as a decrease in

the nuclear content of p27 and its release into the

cytoplasm. An imbalance between the amounts of

nuclear and cytoplasmic p27 is a poor prognostic sign

and is found mainly in malignant skin tumors.

According to the literature (12,13,40,41,46), more than

70% of metastatic melanomas contain p27 in the

cytoplasm, while melanomas without metastases

contain p27 in the nucleus. The appearance of a

significant number of cells with nuclear membrane

staining, predominantly in the irritated and adenoid


background image

Volume 04 Issue 06-2024

57


American Journal Of Biomedical Science & Pharmaceutical Innovation
(ISSN

2771-2753)

VOLUME

04

ISSUE

06

P

AGES

:

54-69

OCLC

1121105677
















































Publisher:

Oscar Publishing Services

Servi

type of KS, indicates a greater risk of malignancy than

in other histological types of tumors, where there is no

membrane staining. A negative reaction during an IHC

study or the detection of single positive nuclei of tumor

cells in hyperkeratotic, acanthotic and clonal (in some

cases) MC can be regarded, on the one hand, as the

absence of significant proliferative activity of cells and

the lack of inclusion of a “reserve” mechanism" in the

form of p27, on the other hand, indicate a defect in

apoptosis, given the slow, steady growth of elements.

In none of these cases, specific staining of nuclear

membranes was detected, which indicates different

mechanisms of cell cycle disorders in different

histological types of keratomas. Thus, morphological

variants of MC have different prognosis for

development, growth and risk of malignant

transformation. The identified features of p27 protein

expression indicate the presence of disturbances in the

regulation of the cell cycle and proliferative activity of

tumor cells, characteristic of each histological type of

KS, which must be taken into account in a

comprehensive assessment of the expression of other

cell cycle markers in KS (13,45).

The role of insulin resistance in the pathogenesis of KS

brings us closer to the theory of keratinocyte aging. In

a study by A. Saraiya et al. in 2013, a case was described

in which the appearance of multiple MCs in the

absence of a genetic predisposition was associated in

patients with insulin resistance and type 2 diabetes

mellitus (type 2 diabetes). Thus, a major role for high

insulin concentrations was suggested in stimulating

DNA synthesis and cell proliferation. Also, M. Blomberg

et al., based on their observations, recommended that

when FGFR mutations are found and concomitant skin

pathology in the form of papillary pigmentary

dystrophy of the skin or multiple MCs, insulin studies

are carried out in order to detect hyperinsulinemia 2. It

should be taken into account that type 2 diabetes is

one of the most common diseases; every 20

inhabitants of the planet suffer from it after 35

40

years. In addition, it is included in the so-called group

of “old age” diseases along with ischemic heart

disease, stroke, and atherosclerosis. In a study by a

number of authors of 150 patients with multiple KS,

type 2 diabetes occurred in 65.3% of cases (98

patients), and impaired glucose tolerance

in 24% of

cases (36 patients). Such a high percentage of a

combination of carbohydrate metabolism disorders

and multiple KS can hardly be considered a mere

coincidence, given that both diseases are genetically

determined and develop in middle and old age.

Interestingly, one of the possible factors in the

development of type 2 diabetes is considered to be an

increase in the level of p16 in the pancreas during

aging, leading to inhibition of beta cell proliferation

and a decrease in their ability to respond to damage,

which subsequently leads to insulin resistance (50).

Morphologically, 6 histological types are distinguished:

acanthotic,

adenoid

(reticular),

hyperkeratotic

(papillomatous),

clonal,

melanoacanthoma

and


background image

Volume 04 Issue 06-2024

58


American Journal Of Biomedical Science & Pharmaceutical Innovation
(ISSN

2771-2753)

VOLUME

04

ISSUE

06

P

AGES

:

54-69

OCLC

1121105677
















































Publisher:

Oscar Publishing Services

Servi

irritated. In all histological types, hyperkeratosis,

acanthosis, papillomatosis, horny and pseudohorny

cysts are present to varying degrees of severity. The

intensity of melanin pigment varies from almost

completely absent to strong. In addition, 2 rare

histological types of seborrheic keratosis have recently

been described - with a large amount of mucin in the

cells - adamantinoid and when basaloid keratinocytes

are arranged like “pseudorosettes”. Currentl

y, there is

no consensus on the etiology and pathogenesis of KS.

Most theories are contradictory and do not explain the

essence of the pathological process and the variety of

forms.

Follicular keratoses represent a disorder of

keratinization and differentiation of keratinocytes,

leading to the formation of keratotic plugs and

parakeratotic cones that penetrate the dermis, causing

perforation of the epidermis. There is no uniform

classification of keratoses pilaris. There are papular,

atrophying and vegetative forms (4,29,30). The

etiology and pathogenesis have not been fully studied.

The previously assumed role of a viral or bacterial

infection, a violation of vitamin A metabolism in the

development of this dermatosis is only of historical

interest. A certain significance in the development of

the disease, in addition to impaired carbohydrate

metabolism, is attributed to liver damage (chronic

hepatitis) with the development of secondary vitamin

A deficiency (4.14). A genetic predisposition to the

development of Kirle disease (KD) has not been

conclusively proven, although cases of the disease

among relatives in the same family with consanguinity

(first-degree relatives) have been described. As

originally defined by J. Kirle (30), it is a disease in which

an atypical clone of keratinocytes penetrates through

the epidermis into the dermis. It is believed that the

basis of the pathological process is a violation of

keratinization, differentiation and keratinization of

keratinocytes (formation of dyskeratotic foci and

acceleration of the keratinization process). This leads

to the formation of keratotic plugs with areas of

parakeratosis. Keratification begins already at the

border of the epidermis and dermis. The rate of

differentiation and keratinization exceeds the rate of

cell proliferation, therefore the parakeratotic cone

partially penetrates deeper into the damaged

epidermis and causes its perforation into the dermis

(4,5,43,49).

There is no uniform classification of keratoses pilaris.

Among the independent nosological forms, papular,

atrophying and vegetative follicular keratoses are

distinguished. Some authors note the similarity of Kirle

disease (KD) with lenticular persistent hyperkeratosis

of Flegel, considering the latter as a variant of KD,

although the clinical manifestations differ. Other

authors (5,7,18) classify CD as vegetative follicular

keratoses, and Flegel's disease as papular.

There is a concept of reactive perforating

keratinization disorder in kidney disease, liver disease

and diabetes mellitus. The concept of perforating skin


background image

Volume 04 Issue 06-2024

59


American Journal Of Biomedical Science & Pharmaceutical Innovation
(ISSN

2771-2753)

VOLUME

04

ISSUE

06

P

AGES

:

54-69

OCLC

1121105677
















































Publisher:

Oscar Publishing Services

Servi

diseases has emerged, in which the elimination of

altered skin components occurs through the epidermis

(transepidermal elimination). Some authors dispute

the independence of CD as a nosological entity (45,47).

In ICD-10, in class XII (diseases of the skin and

subcutaneous tissue), CD has a subcategory L87.0 -

transepidermal perforated changes.

Pathomorphologically, there are depressions in the

epidermis and dilated orifices of hair follicles filled with

hyperkeratotic plugs. Under the plugs, the growth of

the granular layer is pronounced, and in places without

hypergranulosis there is parakeratosis, penetrating to

the dermis. Next, the epidermis becomes thinner and

horny masses penetrate into the dermis, and

inflammatory infiltrates such as granulomas are

formed from lymphocytes, leukocytes, histiocytes and

giant cells. Death of the sebaceous glands,

degeneration of collagen fibers, and hyperelastosis are

observed (49). Clinically, the onset is gradual, new

rashes appear as old ones disappear. Characteristic are

follicular or parafollicular papules, first the color of

healthy skin, then a grayish or brownish-red hue, up to

1 cm in diameter. In the center of the elements there is

a horny plug, when removed, a crater-shaped

depression is formed. Papules tend to grow

peripherally and coalesce, forming dry polycyclic

plaques covered with scales and crusts. The

consistency is dense, the surface is uneven, warty.

Fresh rashes are accompanied by mild itching (more

often in patients with diabetes) or do not bother. Old

lesions are painful when pressed. Localization of

rashes - extensor surfaces of the limbs, torso, buttocks.

Koebner phenomenon and secondary infection are

possible (30). The mucous membranes are not

affected, and rashes on the palms, soles, genitals and

mouth are rare. The course of Kirle disease (KD) is

chronic and relapsing, treatment is difficult, and the

prognosis depends on the underlying disease.

Diagnosis is based on history, clinical and histological

picture. Differential diagnosis includes Devergy's lichen

pilaris, Darier's follicular dyskeratosis, Mibelli's

porokeratosis, elastosis perforating creeping, reactive

perforating collagenosis and Flegel's disease (31,32).

Flegel's disease, like CD, a rare form of keratosis pilaris,

is associated with impaired synthesis of keratin 55K.

Histologically, thinning of the epidermis, follicular

orthokeratosis with parakeratosis, spongiosis and

lymphocytic infiltrate in the dermis are revealed. The

previously assumed importance of Odland bodies in

the pathogenesis of dermatosis is now doubtful. Unlike

CD, this dermatosis appears in adolescence and middle

age and is characterized by small horny papules that

are not prone to plaque formation (27,29,31,32,33).

Actinic keratosis is a skin disease characterized by

limited, dense hyperkeratotic lesions in areas exposed

to solar radiation. The probability of occurrence of

squamous cell carcinoma in lesions of actinic keratosis,

according to some data, is estimated from 0.85 to 10%

per lesion per year [48,49,50]. As a rule, squamous cell

carcinoma that develops in areas of actinic keratosis


background image

Volume 04 Issue 06-2024

60


American Journal Of Biomedical Science & Pharmaceutical Innovation
(ISSN

2771-2753)

VOLUME

04

ISSUE

06

P

AGES

:

54-69

OCLC

1121105677
















































Publisher:

Oscar Publishing Services

Servi

has a favorable course, but in rare cases it can

metastasize (15,16,17).

Porokeratosis is a rare group of acquired or hereditary

dermatoses characterized by linear or annular plaques

with a keratotic border.

Disseminated superficial actinic porokeratosis (DSAP)

is a disease characterized by impaired keratinization.

Disseminated superficial actinic porokeratosis is one of

six types of porokeratosis. It has wider participation

than most other options. These other variants include

linear porokeratosis, Mibelli porokeratosis, punctate

porokeratosis, palmar and plantar disseminated

porokeratosis,

and

disseminated

superficial

porokeratosis. Other rare variants are ptychotropic

porokeratosis,

facial

porokeratosis,

giant

porokeratosis, hypertrophic verrucous porokeratosis,

reticular porokeratosis, and eruptive pruritic papular

porokeratosis. The eruptive form of porokeratosis is

associated with malignancy, immunosuppression, and

a proinflammatory state. Rashes appear all over the

div. A feature that is observed in all these variants is

the horn-shaped plate. On histology, it appears as a

column of parakeratotic cells and is characterized by a

raised ridge bordering the porokeratotic lesions. Risk

factors

for

porokeratosis

include

genetics,

immunosuppression, and ultraviolet light. Lesions in

disseminated superficial actinic porokeratosis begin as

papules and pink to brown macules with a raised

border on exposed skin that may be asymptomatic or

mildly

itchy.

These

lesions

are

considered

precancerous. There is a risk of malignant

transformation to squamous cell or basal cell

carcinoma from 7.5 to 10% (12,22,23)

Eruptive pruritic papular porokeratosis is a rare

subtype of porokeratosis that manifests as an acute

exacerbation of an annular papule with a distinct

peripheral border of a hyperkeratotic ridge and intense

itching. EPPP mainly occurs in older East Asian men. Its

etiology and pathogenesis are unknown (24).

The

development of disseminated superficial

porokeratosis is sometimes observed in association

with renal transplantation, autoimmune diseases, and

various hematologic disorders, suggesting that certain

immunosuppression may cause widespread abnormal

keratinization. It may also be associated with

exacerbation of diabetes mellitus due to the formation

of anti-insulin antibodies (32).

Benign lichenoid keratosis (BLK, LPLK) is clinically

often misdiagnosed as superficial basal cell carcinoma

(BCC), especially when it occurs on the trunk. However,

regression of the BCC may be associated with lichenoid

interface dermatitis, which may be misinterpreted as

BLK on histopathological sections (48).

Benign lichenoid keratosis is a skin lesion consisting of

a non-pruritic papule or slightly indurated plaque,

histologically characterized by a band-shaped

inflammatory infiltrate involving the surface of the

skin.

The authors believe that benign lichenoid keratosis

may be a specific disease rather than an inflammatory


background image

Volume 04 Issue 06-2024

61


American Journal Of Biomedical Science & Pharmaceutical Innovation
(ISSN

2771-2753)

VOLUME

04

ISSUE

06

P

AGES

:

54-69

OCLC

1121105677
















































Publisher:

Oscar Publishing Services

Servi

stage of regressive solar lentigo, large cell acanthoma,

or reticulate seborrheic keratosis (16,17,19).

A study of palmoplantar keratoderma in eighty-two

cases showed that the cause of palmoplantar

keratoderma is twenty different diseases, both

hereditary and acquired. The maximum number of

cases occurs in the hereditary variety of palmoplantar

keratoderma (Unna-Tost syndrome) (28.05%). While

among acquired diseases the leading cause was

psoriasis (17.07%). Two histopathological types of

Unna-Tost syndrome and their correlation with clinical

features have been reported (4,5)

Keratoderma spinosa is a rare dermatosis consisting of

multiple projections located on the palms and soles,

with a distinct histopathological feature of a

parakeratotic column over a hypogranular epidermis.

Although there are some hereditary cases, most are

acquired. The latter may be idiopathic or associated

with neoplasms and chronic systemic diseases (15,25).

In some literature sources, keratoderma spinosum is

referred to as keratoderma spinosa. This condition is

identified under several names, such as “music box

spinal

keratosis”

and

“palmoplantar

filiform

hyperkeratosis,” which creates ambiguity in the

diagnostic and histopathological features of the

disease. Because of its association with cancer, all

patients with keratoderma spinosa should undergo

baseline cancer screening based on age and then once

or twice a year or as symptoms appear (3,4).

The incidence of skin malignancies has been increasing

over the years (46,47). Since 1960, their incidence

worldwide has increased annually by 4

8% and

amounts to about 3 million newly identified cases per

year. One of the most significant etiological factors is

considered to be excessive insolation - frequent

exposure to the sun for a long time and repeated

sunburn. In many cases, malignant epithelial skin

tumors develop in the setting of preexisting

dermatoses such as actinic keratosis and Bowen's

disease (22).

Bowen's disease is an intraepidermal form of

squamous cell carcinoma (or squamous cell carcinoma

in situ) in the form of a single, slowly growing plaque.

Its development is most often associated with

exposure to ultraviolet radiation, less often with skin

trauma or contact with arsenic. In this regard, two

forms of the disease are considered: one is localized in

open areas of the skin (exposed to insolation), and the

other is localized in closed areas. The course of

Bowen's disease is steadily progressive, although in

the vast majority of cases it remains cancer in situ

throughout the patient's life. Invasive squamous cell

skin cancer develops against the background of

Bowen's disease with a frequency of 5% to 11% of cases

of long-term existence of the pathology. As long as the

disease remains in the intraepidermal stage,

metastases do not occur. It is possible to differentiate

actinic keratosis and Bowen's disease only on the basis

of histological examination. Thus, with actinic


background image

Volume 04 Issue 06-2024

62


American Journal Of Biomedical Science & Pharmaceutical Innovation
(ISSN

2771-2753)

VOLUME

04

ISSUE

06

P

AGES

:

54-69

OCLC

1121105677
















































Publisher:

Oscar Publishing Services

Servi

keratosis, dysplastic changes in the epidermis are

observed, which occupy no more than

of its

thickness, sometimes with penetration of epidermal

strands into the upper parts of the dermis. However,

atypical cells can spread throughout the entire

thickness of the epidermis - in such cases they are

called bowenoid actinic keratosis (22,24). Bowen's

disease is characterized by a sharp thickening of the

epidermis, consisting of enlarged cells, in places piling

up on each other, with pronounced polymorphism and

polychromasia, with acanthotic growths to the

reticular layer of the dermis (23,26). Proliferation of

atypical keratinocytes, as in bowenoid actinic

keratosis, is observed throughout the entire thickness

of the epidermis. Actinic keratosis with foci of

Bowenization and Bowen's disease are distinguished

by dysplastic changes in the keratinocytes of the

follicular epithelium - a sign inherent in Bowen's

disease. In addition, with bowenoid actinic keratosis,

manifestations of solar elastosis are always observed

in the dermis. However, according to some authors,

the bowenoid type of solar keratosis is histologically

indistinguishable from Bowen's disease and at this

stage of development is regarded as carcinoma in situ

(1,22,6,28). Despite the atypia of keratinocytes,

characteristic of both actinic keratosis and Bowen's

disease, there are no signs of true invasion - the

boundary between the epidermis and dermis remains

clear.

Of particular importance in all cases is the study of the

expression of a number of cellular markers involved in

the mechanisms of development of malignant skin

tumors, such as oncoproteins associated with

intercellular adhesion and proliferation. Several

nuclear and membrane antigens are known, changes in

expression of which are due to proliferative activity

cells. One of the most studied molecular biomarkers is

the indicator of proliferative activity Ki-67, whose

antibodies react with proliferating cells. If the cell does

not proliferate, this interaction does not occur.

Normally, in healthy epidermis, Ki-67 expression is

observed only in the basal layer, and the average

proliferative index, according to various sources, varies

from 0.8 to 11% (24,25,34,35,36).

The architectural integrity of the epidermis is ensured

by keratins and intercellular adhesion molecules -

epithelial cadherins. E-cadherin is a calcium-dependent

adhesion molecule characteristic of epithelial tissue

cells. Its long extracellular sections form parallel

dimers on the cell surface, which, when in contact with

E-cadherin molecules of neighboring keratinocytes,

form strong zipper-type bonds. A decrease in

intercellular adhesion allows affected cells to split off

from normal ones, which leads to destruction of

histological structures. Normally, in the epidermis, E-

cadherin is detected in 100% of cells in the form of

uniform

membrane

staining.

Increased

cell

proliferative activity and impaired expression of E-


background image

Volume 04 Issue 06-2024

63


American Journal Of Biomedical Science & Pharmaceutical Innovation
(ISSN

2771-2753)

VOLUME

04

ISSUE

06

P

AGES

:

54-69

OCLC

1121105677
















































Publisher:

Oscar Publishing Services

Servi

cadherin indicate an increase in cell invasive capacity

(26,27).

Data from a number of authors indicate that actinic

keratosis, bowenoid actinic keratosis and Bowen's

disease represent different stages of the development

of the same malignant process. The predominance of

low proliferative activity, the expression of Ki-67 in the

lower parts of the epidermis and the preservation of

adhesive interactions between cells in actinic keratosis

indicate the initial stages of the process and its low

invasive potential (22,23).

Epithelial skin tumors are common human neoplasms,

accounting for from 20

24.9 to 55.4

61.7% of calls for

skin neoplasms. One of the key factors contributing to

the activation of proliferative processes in the skin with

the subsequent development of neoplasms of various

natures is the infection of skin cells with the human

papillomavirus (HPV). Certain characteristic HPV

fragments suppress the activity of the keratinocyte p53

gene, which leads to uncontrolled proliferation of

keratinocytes. The importance of certain types of HPV

in the development of a number of skin tumors has

been shown (27,47,49).

The PCNA protein is recognized as a more accurate

indicator of proliferation, identifying cells in the

process of preparing for division in the S-phase of the

cell cycle. The evidence that the S-phase in cells

affected by HPV is longer (from 18 to 20 hours) than the

S-phase in normal epidermal keratinocytes (16 hours) is

confirmed

by

immunohistochemical

studies.

Seborrheic keratosis is the most common benign skin

tumor in different regions of the world (47,49).

It is known that the vast majority of proto-oncogenes

and tumor suppressors are components of several

common signaling pathways that control the cell cycle,

apoptosis, genome integrity, morphogenetic reactions

and cell differentiation. Changes in these signaling

pathways ultimately lead to the development of

tumors. To date, about a hundred potential oncogenes

(cellular and viral) and about two dozen tumor

suppressors are known. Changes characteristic of

predominantly malignant human tumors have been

identified, including highly specific anomalies used to

make a diagnosis, while almost no attention has been

paid to benign formations (16,22).

Two families of proteins, inhibitors of cyclin-dependent

kinases, have been identified: Ink4 and Cip/Kip. The

first includes four members, including the tumor

suppressors p15INK4b and p16INK4a. Ink4 proteins

have a fairly narrow specificity: by binding the cyclin-

dependent kinase (Cdk) Cdk4 and Cdk6, they prevent

the formation of their complexes with cyclins D. The

Cip/Kip

family

consists

of

three

members:

p21WAF1/CIP1, p27KIP1a and p57KIP2. These proteins

bind and inhibit already fully formed complexes of

cyclin D

Cdk4(6), cyclin E

Cdk2 and cyclin A

Cdk2

[5, 6]. Inhibition of the functions of cyclin-dependent

kinases leads to hypophosphorylation of the pRB

protein, which reduces the expression of E2F-

dependent genes, blocks the transition of the cell from


background image

Volume 04 Issue 06-2024

64


American Journal Of Biomedical Science & Pharmaceutical Innovation
(ISSN

2771-2753)

VOLUME

04

ISSUE

06

P

AGES

:

54-69

OCLC

1121105677
















































Publisher:

Oscar Publishing Services

Servi

the G1 phase to the S phase, and controls cell division

and proliferation. Many sources confirm that

disturbances in the G1 phase and G1/S checkpoint lead

to uncontrolled tumor growth. Thus, it has been

experimentally proven that a decrease in p16 protein

expression leads to hyperphosphorylation of pRB and

subsequently leads to activation of the transcription of

S-phase-specific genes. The p16 protein was discovered

by researchers in 1993 and since its discovery has

become one of the most sought-after markers in the

field of cancer research [10]. It is encoded by the tumor

suppressor gene CDKN2A, located on chromosome 9

(9p21.3). The participation of this gene has been noted

in the development of both sporadic and familial forms

of melanoma, glioma, lung cancer, T-cell leukemia, and

B-cell leukemia. In addition, p16 is currently used as a

prognostic biomarker for patients with oropharyngeal

squamous cell carcinoma and cervical cancer. The

expression of this protein and its role in the

pathogenesis of benign skin tumors, in particular

seborrheic keratosis, have been little studied

(22,24,44,45,49).

Thus, Y. H. Wu et al. in 17 cases of seborrheic keratosis

with signs of bowenoid transformation, he found

increased expression of p16 and p21 in the cells, which

is also characteristic of Bowen's disease and bowenoid

papulosis. They were asked to study p16 in patients

with seborrheic keratosis to identify possible

malignancy of the elements. Other authors - S.

Nakamura et al., having identified in skin samples from

KS lesions (including in cultured keratinocytes from

patients with KS) a pronounced expression of the p16

protein in all tumor cells, associated this not with the

possibility of malignancy, but with a violation of the cell

cycle , blocking the entry into the S-phase of cells and

their premature aging. In support of this hypothesis, in

the 4 studied SC samples, genetic analysis revealed the

absence of DNA fragmentation in tumor cells, whereas

in normal epidermis fragmentation was present in the

granular and stratum corneum. This indicates that

apoptosis is inhibited in KS. Brueks et al., studying

10 cases of seborrheic keratomas of exclusively

acanthotic and irritable types, noted an average level

of expression of the p16 protein and a pronounced,

diffuse expression of another protein, the p27 kinase

inhibitor, which, in his opinion, indicates the dominant

influence of this protein on cell proliferative activity.

Like p16, the p27 protein (Kip1) regulates the course of

the cell cycle and is responsible for its arrest in the G1

phase. It is the product of the human CDKN1B gene

(22,43).

So, today, ideas about the role of proteins - inhibitors

of cyclin-dependent kinases in oncogenesis are not

entirely clear; the data from most studies are

contradictory. The possibility of using them in practice

as prognostic biomarkers opens up prospects for an

individual approach to the treatment of each patient,

understanding the pathogenesis of many both

malignant and benign human tumors, which


background image

Volume 04 Issue 06-2024

65


American Journal Of Biomedical Science & Pharmaceutical Innovation
(ISSN

2771-2753)

VOLUME

04

ISSUE

06

P

AGES

:

54-69

OCLC

1121105677
















































Publisher:

Oscar Publishing Services

Servi

determines the relevance and need for their further

study.

CONCLUSIONS

Studying the morphological and genetic different

types of keratoses is key for correct diagnosis and

choice of treatment. The association between

metabolic disorders and skin diseases highlights the

need for an integrated approach to the management

of patients with chronic diseases, including diabetes

mellitus. Further research is needed to better

understand the pathogenesis and develop effective

treatments.

REFERENCES

1.

Кожные

и

венерические

болезни.

Справочник. Под ред. О.Л.Иванова. М:

Медицина 2007; 117—

118.

2.

Родионов А. Н., Заславский Д. В., Сыдиков А.

А. Клиническая дерматология. –

2019.

3.

Болезнь Кирле (кератоз фолликулярный и

парафолликулярный,

проникающий):

клинический

случай

В.В.

КАРПОВ1КЛИНИЧЕСКАЯ ДЕРМАТОЛОГИЯ И

ВЕНЕРОЛОГИЯ 5, 2011стр.25

-28).

4.

Huang J. et al. An epidemiological study on skin

tumors of the elderly in a community in

Shanghai, China //Scientific Reports.

2023.

Т.

13.

№. 1. –

С. 4441.

5.

Alapatt G. F., Sukumar D., Bhat M. R. A

clinicopathological

and

dermoscopic

correlation of seborrheic keratosis //Indian

Journal of Dermatology.

2016.

Т. 61. –

№. 6.

С. 622

-627.

6.

Kim H.S., Park E.J., Kwon I.H., Kim K.H., Kim K.J.

Clinical and histopathologic study of benign

lichenoid keratosis on the face. Am. J.

Dermatopathol. 2013; 35(7): 738

41.].

7.

Mohamed M., Amri M., Njim L., Jribi M.,

Zakhama A., Zili J. Pigmented keratosis on the

face. Ann. Dermatol. Venereol. 2013; 140(5):

390-2.

8.

Bruecks

A.K.,

Kalia

S.,

Trotter

M.J.

Overexpression of p27 KIP1 in seborrheic

keratosis. J. Cutan. Med. Surg. 2007; 11(5): 174

8.

9.

Hafner C., van Oers J.M., Hartmann A.,

Landthaler M., Stoehr R., Blaszyk H., et al. High

frequency of FGFR3 mutations in ad¬enoid

seborrheic keratoses. J. Invest. Dermatol.

2006; 126(11): 2404

7.,

10.

Ruas M., Gregory .F, Jones R., Poolman R.,

Starborg M., Rowe J., et al. CDK4 and CDK6

delay senescence by kinase-dependent and

p16INK4a-independent mechanisms. Mol. Cell

Biol. 2007; 27(12): 4273

82.

11.

Seargent J.M., Loadman P.M., Martin S.W.,

Naylor B., Bibby M.C., Gill J.H. Expression of

matrix metalloproteinase-10 in human bladder

transitional cell carcinoma. Urology. 2005;

65(4): 815-20.


background image

Volume 04 Issue 06-2024

66


American Journal Of Biomedical Science & Pharmaceutical Innovation
(ISSN

2771-2753)

VOLUME

04

ISSUE

06

P

AGES

:

54-69

OCLC

1121105677
















































Publisher:

Oscar Publishing Services

Servi

12.

Morgan D. Cell Cycle: Principals of Control.

London: New Sci¬ence Press Ltd.; 2007: 145

6.,

Mitrea D.M., Yoon M.K., Ou L., Kriwacki R.W.

Disorder-func¬tion relationships for the cell

cycle regulatory proteins p21 and p27. Biol.

Chem. 2012; 393(4): 259

74.

13.

Mitrea D.M., Yoon M.K., Ou L., Kriwacki R.W.

Disorder-func¬tion relationships for the cell

cycle regulatory proteins p21 and p27. Biol.

Chem. 2012; 393(4): 259

74.

14.

Pellegata

N.S.,

Quintanilla-Martinez

L.,

Siggelkow H., Sam¬son E., Bink K., Hofler H., et

al. Germ-line mutations in p27 Kip1 cause a

multiple endocrine neoplasia syndrome in rats

and humans. Proc. Natl. Acad. Sci. U.S.A. 2006;

103(42): 15558

63.

15.

Александрова

А.К.1,

Суколин

Г.И.2,

Смольянникова

В.А.ОСОБЕННОСТИ

ЭКСПРЕССИИ БЕЛКА p27 ПРИ РАЗНЫХ

ФОРМАХ

СЕБОРЕЙНОГО

КЕРАТОЗА

Российский журнал кожных и венерических

болезней 2016,19 (5) стр.283

-286.

16.

Karadag A. S., Parish L. C. The status of the

seborrheic keratosis //Clinics in Dermatology.

2018.

Т. 36. –

№. 2. –

С. 275

-277.

17.

Jackson J. M. et al. Current understanding of

seborrheic keratosis: prevalence, etiology,

clinical

presentation,

diagnosis,

and

management

//Journal

of

drugs

in

dermatology: JDD.

2015.

Т. 14. –

№. 10. –

С.

1119-1125.

18.

Sanders M. G. H. et al. Prevalence and

determinants of seborrhoeic dermatitis in a

middle‐aged and elderly population: the

Rotterdam

Study

//British

Journal

of

Dermatology.

2018.

Т. 178. –

№. 1. –

С. 148

-

153.

19.

Squillace L. et al. Unusual dermoscopic

patterns

of

seborrheic

keratosis

//Dermatology.

2016.

Т. 232. –

№. 2. –

С. 198

-

202.

20.

Wollina U. Recent advances in managing and

understanding

seborrheic

keratosis

//F1000Research.

2019.

Т. 8.

21.

Del Rosso J. Q. A closer look at seborrheic

keratoses:

patient

perspectives,

clinical

relevance, medical necessity, and implications

for management //The Journal of clinical and

aesthetic dermatology.

2017.

Т. 10. –

№. 3. –

С. 16.

22.

Bagazgoitia L, Cuevas J, Juarranz A. Expression

of p53 and p16 in actinic keratosis, Bowenoid

actinic keratosis and Bowen's disease. J Eur

Acad Dermatol Venereol. 2010;24(2):228

30.

23.

Broers J. L. V., Ramaekers F. C. S. The role of the

nuclear lamina in cancer and apoptosis //Cancer

Biology and the Nuclear Envelope: Recent

Advances May Elucidate Past Paradoxes.

2014.

С. 27

-48.


background image

Volume 04 Issue 06-2024

67


American Journal Of Biomedical Science & Pharmaceutical Innovation
(ISSN

2771-2753)

VOLUME

04

ISSUE

06

P

AGES

:

54-69

OCLC

1121105677
















































Publisher:

Oscar Publishing Services

Servi

24.

Talghini S, Halimi M, Baybordi H. Expression of

p 27, Ki 67, p 53 in squamous cell carcinoma,

actinic keratosis and Bowen disease. Pakistan

Journal of Biological Sciences. 2009;12(12):929

33. doi: 10.3923/pjbs.2009.929.933.

25.

Ahlgrimm-Siess V. et al. Seborrheic keratosis:

reflectance confocal microscopy features and

correlation with dermoscopy //Journal of the

American Academy of Dermatology.

2013.

Т.

69.

№. 1. –

С. 120

-126.

26.

Papadavid E, Pignatelli M, Zakynthinos S,

Krausz T, Chu AC. Abnormal immunoreactivity

of the E-cadherin/catenin (alpha-, beta-, and

gamma-) complex in premalignant and

malignant non-melanocytic skin tumours. J

Pathol.2002;196(2):154

62.

27.

Fuller LC, Allen MH, Montesu M, Barker JN,

Macdonald DM. Expression of E-cadherin in

human epidermal non-melanoma cutaneous

tumours. Br J Dermatol. 1996;134(1):28

32.).

28.

Хлебникова А.Н., Гуревич Л.Е.,

Селезнева

Е.В., Обыденова К.В., Седова Т.Г.,Кунцевич

Ж.С.

Особенности

пролиферации

и

межклеточной адгезии в предраковых и

злокачественных эпителиальных опухолях

кожи. Альманах клинической медицины.

2016 Январь; 44 (1): 58–

63.

29.

Aljada A. et al. Altered Lamin A/C splice variant

expression as a possible diagnostic marker in

breast cancer //Cellular Oncology.

2016.

Т.

39.

С. 161

-174.

30.

Dubik N., Mai S. Lamin A/C: function in normal

and tumor cells //Cancers.

2020.

Т. 12. –

№.

12.

С. 3688.

31.

Blomberg M., Jeppesen E.M., Skovby F.,

Benfeldt E. "FGFR3 Mutations and the Skin:

Report of a Patient with a FGFR3 Gene

Mutation,

Acanthosis

Nigricans,

Hypochondroplasia and Hyperinsulinemia and

Review of the Literature." Dermatology 220.4

(2010):297

300. DOI: 10.1159/000297575.

32.

Kim Y. The impact of altered lamin B1 levels on

nuclear lamina structure and function in aging

and human diseases //Current Opinion in Cell

Biology.

2023.

Т. 85. –

С. 102257.

33.

Saraiya A., Al-Shoha A., Brodell R.T.

"Hyperinsulinemia Associated with Acanthosis

Nigricans, Finger Pebbles, Acrochordons, and

the Sign of Leser-Trélat." Endocr. Pract. 19.3

(2013): 522

555. DOI: 10.4158/EP12192.RA.

34.

Squillace L, Cappello M, Longo C, Moscarella E,

Alfano R, Argenziano G. Unusual dermoscopic

patterns of seborrheic keratosis. Dermatology.

2016;232(2): 198

202. doi:10.1159/000442439.

19.

35.

Roh NK, Hahn HJ, Lee YW, Choe YB, Ahn KJ.

Clinical and histopathological investigation of

seborrheic

keratosis.

Ann

Dermatol.

2016;28(2):152

8. doi: 10.5021/ad.2016.28.2.152.


background image

Volume 04 Issue 06-2024

68


American Journal Of Biomedical Science & Pharmaceutical Innovation
(ISSN

2771-2753)

VOLUME

04

ISSUE

06

P

AGES

:

54-69

OCLC

1121105677
















































Publisher:

Oscar Publishing Services

Servi

36.

Tardío J.C., Bancalari E., Moreno A., Martín-

Fragueiro L.M. "Genital Seborrheic Keratoses

Are Human Papillomavirus-related Lesions. A

Linear Array Genotyping Test Study." APMIS.

120.6 (2012): 477

483. DOI: 10.1111/j.1600-

0463.2011.02853.x.,

37.

Jackson JM, Alexis A, Berman B, Berson

DS,Taylor S, Weiss JS. Current understanding

of seborrheic keratosis: prevalence, etiology,

clinical

presentation,

diagnosis,

and

management.J Drugs Dermatol. 2015;14(10):

1119

25. 18.

38.

Squillace L, Cappello M, Longo C, Moscarella E,

Alfano R, Argenziano G. Unusual dermoscopic

patterns of seborrheic keratosis. Dermatology.

2016;232(2): 198

202. doi: 10.1159/000442439.

19.

39.

Roh NK, Hahn HJ, Lee YW, Choe YB, Ahn KJ.

Clinicaland histopathological investigation of

seborrheic

keratosis.

Ann

Dermatol.

2016;28(2): 152

8. doi: 10.5021/ad.2016.28.2.152.

40.

Martin B.C., Warram J.H., Krolewski A.S.,

Bergman R.N., Soeldner J.S., Kahn C.R. "Role of

Glucose and Insulin Resistance in Development

of Type 2 Diabetes Mellitus: Results of a 25-year

Follow-up Study." Lancet 340 (1992): 925

929.,

41.

Юсупова Л. А. и др. Патогенетические

механизмы, диагностика и клиническая

картина себорейного кератоза //Лечащий

врач. –

2018.

№. 10. –

С. 73.

42.

Saraiya A., Al-Shoha A., Brodell R.T.

"Hyperinsulinemia Associated with Acanthosis

Nigricans, Finger Pebbles, Acrochordons, and

the Sign of Leser-Trélat." Endocr. Pract. 19.3

(2013): 522

555.

43.

Elder D.E., Elenitsas R., Johnson B.L. Jr, Murphy

G.F., Xu X. Lever's Histopathology of the Skin.

Philadelphia: Lippincott-Raven, 2009. 795

798

44.

Пашинян А. Г., Джаваева Д. Г., Молчанова О.

В. Себорейный дерматит в практике врача

-

косметолога.

Клинические

проявления.

Диагностика. Лечение //Метаморфозы. –

2018.

№. 24. –

С. 12

-16..

45.

Alapatt GF, Sukumar D, Bhat MR. A

clinicopathological

and

dermoscopic

correlation of seborrheic keratosis. Indian J

Dermatol. 2016;61(6):622

7.

46.

Saraiya A., Al-Shoha A., Brodell R.T.

"Hyperinsulinemia Associated with Acanthosis

Nigricans, Finger Pebbles, Acrochordons, and

the Sign of Leser-Trélat." Endocr. Pract. 19.3

(2013): 522

555. DOI: 10.4158/EP12192.RA.

47.

Choi HJ, Lee JH. Multiple human papilloma

virus 16 infection presenting as various skin

lesions. J Craniofac Surg. 2016;27(4):e379

81.

48.

Wu YH, Hsiao PF, Chen CK. Seborrheic keratosis

with

bowenoid

transformation:

the

immunohistochemical

features

and

its

association

with

human

papillomavirus


background image

Volume 04 Issue 06-2024

69


American Journal Of Biomedical Science & Pharmaceutical Innovation
(ISSN

2771-2753)

VOLUME

04

ISSUE

06

P

AGES

:

54-69

OCLC

1121105677
















































Publisher:

Oscar Publishing Services

Servi

infection. Am J Dermatopathol. 2015;37(6):

462

8.

49.

Писклакова Т.П., Костенко Е.И., Телешева

Л.Ф. Себорейный кератоз: клинические

особенности и ассоциация с вирусом

папилломы человека рода β Альманах

клинической медицины. 2017 Март

-

апрель;

45 (2): 118

126.

50.

Rayess H., Wang M. B., Srivatsan E. S. Cellular

senescence and tumor suppressor gene p16 //

Int. j. cancer

2012.

V. 130 (8).

P. 171

174.,

References

Кожные и венерические болезни. Справочник. Под ред. О.Л.Иванова. М: Медицина 2007; 117—118.

Родионов А. Н., Заславский Д. В., Сыдиков А. А. Клиническая дерматология. – 2019.

Болезнь Кирле (кератоз фолликулярный и парафолликулярный, проникающий): клинический случай В.В. КАРПОВ1КЛИНИЧЕСКАЯ ДЕРМАТОЛОГИЯ И ВЕНЕРОЛОГИЯ 5, 2011стр.25-28).

Huang J. et al. An epidemiological study on skin tumors of the elderly in a community in Shanghai, China //Scientific Reports. – 2023. – Т. 13. – №. 1. – С. 4441.

Alapatt G. F., Sukumar D., Bhat M. R. A clinicopathological and dermoscopic correlation of seborrheic keratosis //Indian Journal of Dermatology. – 2016. – Т. 61. – №. 6. – С. 622-627.

Kim H.S., Park E.J., Kwon I.H., Kim K.H., Kim K.J. Clinical and histopathologic study of benign lichenoid keratosis on the face. Am. J. Dermatopathol. 2013; 35(7): 738–41.].

Mohamed M., Amri M., Njim L., Jribi M., Zakhama A., Zili J. Pigmented keratosis on the face. Ann. Dermatol. Venereol. 2013; 140(5): 390-2.

Bruecks A.K., Kalia S., Trotter M.J. Overexpression of p27 KIP1 in seborrheic keratosis. J. Cutan. Med. Surg. 2007; 11(5): 174–8.

Hafner C., van Oers J.M., Hartmann A., Landthaler M., Stoehr R., Blaszyk H., et al. High frequency of FGFR3 mutations in ad¬enoid seborrheic keratoses. J. Invest. Dermatol. 2006; 126(11): 2404–7.,

Ruas M., Gregory .F, Jones R., Poolman R., Starborg M., Rowe J., et al. CDK4 and CDK6 delay senescence by kinase-dependent and p16INK4a-independent mechanisms. Mol. Cell Biol. 2007; 27(12): 4273–82.

Seargent J.M., Loadman P.M., Martin S.W., Naylor B., Bibby M.C., Gill J.H. Expression of matrix metalloproteinase-10 in human bladder transitional cell carcinoma. Urology. 2005; 65(4): 815-20.

Morgan D. Cell Cycle: Principals of Control. London: New Sci¬ence Press Ltd.; 2007: 145–6., Mitrea D.M., Yoon M.K., Ou L., Kriwacki R.W. Disorder-func¬tion relationships for the cell cycle regulatory proteins p21 and p27. Biol. Chem. 2012; 393(4): 259–74.

Mitrea D.M., Yoon M.K., Ou L., Kriwacki R.W. Disorder-func¬tion relationships for the cell cycle regulatory proteins p21 and p27. Biol. Chem. 2012; 393(4): 259–74.

Pellegata N.S., Quintanilla-Martinez L., Siggelkow H., Sam¬son E., Bink K., Hofler H., et al. Germ-line mutations in p27 Kip1 cause a multiple endocrine neoplasia syndrome in rats and humans. Proc. Natl. Acad. Sci. U.S.A. 2006; 103(42): 15558–63.

Александрова А.К.1, Суколин Г.И.2, Смольянникова В.А.ОСОБЕННОСТИ ЭКСПРЕССИИ БЕЛКА p27 ПРИ РАЗНЫХ ФОРМАХ СЕБОРЕЙНОГО КЕРАТОЗА Российский журнал кожных и венерических болезней 2016,19 (5) стр.283-286.

Karadag A. S., Parish L. C. The status of the seborrheic keratosis //Clinics in Dermatology. – 2018. – Т. 36. – №. 2. – С. 275-277.

Jackson J. M. et al. Current understanding of seborrheic keratosis: prevalence, etiology, clinical presentation, diagnosis, and management //Journal of drugs in dermatology: JDD. – 2015. – Т. 14. – №. 10. – С. 1119-1125.

Sanders M. G. H. et al. Prevalence and determinants of seborrhoeic dermatitis in a middle‐aged and elderly population: the Rotterdam Study //British Journal of Dermatology. – 2018. – Т. 178. – №. 1. – С. 148-153.

Squillace L. et al. Unusual dermoscopic patterns of seborrheic keratosis //Dermatology. – 2016. – Т. 232. – №. 2. – С. 198-202.

Wollina U. Recent advances in managing and understanding seborrheic keratosis //F1000Research. – 2019. – Т. 8.

Del Rosso J. Q. A closer look at seborrheic keratoses: patient perspectives, clinical relevance, medical necessity, and implications for management //The Journal of clinical and aesthetic dermatology. – 2017. – Т. 10. – №. 3. – С. 16.

Bagazgoitia L, Cuevas J, Juarranz A. Expression of p53 and p16 in actinic keratosis, Bowenoid actinic keratosis and Bowen's disease. J Eur Acad Dermatol Venereol. 2010;24(2):228–30.

Broers J. L. V., Ramaekers F. C. S. The role of the nuclear lamina in cancer and apoptosis //Cancer Biology and the Nuclear Envelope: Recent Advances May Elucidate Past Paradoxes. – 2014. – С. 27-48.

Talghini S, Halimi M, Baybordi H. Expression of p 27, Ki 67, p 53 in squamous cell carcinoma, actinic keratosis and Bowen disease. Pakistan Journal of Biological Sciences. 2009;12(12):929–33. doi: 10.3923/pjbs.2009.929.933.

Ahlgrimm-Siess V. et al. Seborrheic keratosis: reflectance confocal microscopy features and correlation with dermoscopy //Journal of the American Academy of Dermatology. – 2013. – Т. 69. – №. 1. – С. 120-126.

Papadavid E, Pignatelli M, Zakynthinos S, Krausz T, Chu AC. Abnormal immunoreactivity of the E-cadherin/catenin (alpha-, beta-, and gamma-) complex in premalignant and malignant non-melanocytic skin tumours. J Pathol.2002;196(2):154–62.

Fuller LC, Allen MH, Montesu M, Barker JN, Macdonald DM. Expression of E-cadherin in human epidermal non-melanoma cutaneous tumours. Br J Dermatol. 1996;134(1):28–32.).

Хлебникова А.Н., Гуревич Л.Е., Селезнева Е.В., Обыденова К.В., Седова Т.Г.,Кунцевич Ж.С. Особенности пролиферации и межклеточной адгезии в предраковых и злокачественных эпителиальных опухолях кожи. Альманах клинической медицины. 2016 Январь; 44 (1): 58–63.

Aljada A. et al. Altered Lamin A/C splice variant expression as a possible diagnostic marker in breast cancer //Cellular Oncology. – 2016. – Т. 39. – С. 161-174.

Dubik N., Mai S. Lamin A/C: function in normal and tumor cells //Cancers. – 2020. – Т. 12. – №. 12. – С. 3688.

Blomberg M., Jeppesen E.M., Skovby F., Benfeldt E. "FGFR3 Mutations and the Skin: Report of a Patient with a FGFR3 Gene Mutation, Acanthosis Nigricans, Hypochondroplasia and Hyperinsulinemia and Review of the Literature." Dermatology 220.4 (2010):297–300. DOI: 10.1159/000297575.

Kim Y. The impact of altered lamin B1 levels on nuclear lamina structure and function in aging and human diseases //Current Opinion in Cell Biology. – 2023. – Т. 85. – С. 102257.

Saraiya A., Al-Shoha A., Brodell R.T. "Hyperinsulinemia Associated with Acanthosis Nigricans, Finger Pebbles, Acrochordons, and the Sign of Leser-Trélat." Endocr. Pract. 19.3 (2013): 522–555. DOI: 10.4158/EP12192.RA.

Squillace L, Cappello M, Longo C, Moscarella E, Alfano R, Argenziano G. Unusual dermoscopic patterns of seborrheic keratosis. Dermatology. 2016;232(2): 198–202. doi:10.1159/000442439. 19.

Roh NK, Hahn HJ, Lee YW, Choe YB, Ahn KJ. Clinical and histopathological investigation of seborrheic keratosis. Ann Dermatol. 2016;28(2):152–8. doi: 10.5021/ad.2016.28.2.152.

Tardío J.C., Bancalari E., Moreno A., Martín-Fragueiro L.M. "Genital Seborrheic Keratoses Are Human Papillomavirus-related Lesions. A Linear Array Genotyping Test Study." APMIS. 120.6 (2012): 477–483. DOI: 10.1111/j.1600-0463.2011.02853.x.,

Jackson JM, Alexis A, Berman B, Berson DS,Taylor S, Weiss JS. Current understanding of seborrheic keratosis: prevalence, etiology, clinical presentation, diagnosis, and management.J Drugs Dermatol. 2015;14(10): 1119–25. 18.

Squillace L, Cappello M, Longo C, Moscarella E, Alfano R, Argenziano G. Unusual dermoscopic patterns of seborrheic keratosis. Dermatology. 2016;232(2): 198–202. doi: 10.1159/000442439. 19.

Roh NK, Hahn HJ, Lee YW, Choe YB, Ahn KJ. Clinicaland histopathological investigation of seborrheic keratosis. Ann Dermatol. 2016;28(2): 152–8. doi: 10.5021/ad.2016.28.2.152.

Martin B.C., Warram J.H., Krolewski A.S., Bergman R.N., Soeldner J.S., Kahn C.R. "Role of Glucose and Insulin Resistance in Development of Type 2 Diabetes Mellitus: Results of a 25-year Follow-up Study." Lancet 340 (1992): 925–929.,

Юсупова Л. А. и др. Патогенетические механизмы, диагностика и клиническая картина себорейного кератоза //Лечащий врач. – 2018. – №. 10. – С. 73.

Saraiya A., Al-Shoha A., Brodell R.T. "Hyperinsulinemia Associated with Acanthosis Nigricans, Finger Pebbles, Acrochordons, and the Sign of Leser-Trélat." Endocr. Pract. 19.3 (2013): 522–555.

Elder D.E., Elenitsas R., Johnson B.L. Jr, Murphy G.F., Xu X. Lever's Histopathology of the Skin. Philadelphia: Lippincott-Raven, 2009. 795–798

Пашинян А. Г., Джаваева Д. Г., Молчанова О. В. Себорейный дерматит в практике врача-косметолога. Клинические проявления. Диагностика. Лечение //Метаморфозы. – 2018. – №. 24. – С. 12-16..

Alapatt GF, Sukumar D, Bhat MR. A clinicopathological and dermoscopic correlation of seborrheic keratosis. Indian J Dermatol. 2016;61(6):622–7.

Saraiya A., Al-Shoha A., Brodell R.T. "Hyperinsulinemia Associated with Acanthosis Nigricans, Finger Pebbles, Acrochordons, and the Sign of Leser-Trélat." Endocr. Pract. 19.3 (2013): 522–555. DOI: 10.4158/EP12192.RA.

Choi HJ, Lee JH. Multiple human papilloma virus 16 infection presenting as various skin lesions. J Craniofac Surg. 2016;27(4):e379–81.

Wu YH, Hsiao PF, Chen CK. Seborrheic keratosis with bowenoid transformation: the immunohistochemical features and its association with human papillomavirus infection. Am J Dermatopathol. 2015;37(6): 462–8.

Писклакова Т.П., Костенко Е.И., Телешева Л.Ф. Себорейный кератоз: клинические особенности и ассоциация с вирусом папилломы человека рода β Альманах клинической медицины. 2017 Март-апрель; 45 (2): 118–126.

Rayess H., Wang M. B., Srivatsan E. S. Cellular senescence and tumor suppressor gene p16 // Int. j. cancer – 2012. – V. 130 (8). – P. 171–174.,