Authors

  • Khamidova Farida Muinovna
    Samarkand State Medical University, Department of Pathological Anatomy with a sectional course, Uzbekistan
  • Khusinova Firuza Azgar Kizi
    Samarkand State Medical University, Department of Pathological Anatomy with a sectional course, Uzbekistan

DOI:

https://doi.org/10.37547/ajbspi/Volume04Issue07-05

Keywords:

Acne molecular genetic mechanisms prognosis

Abstract

Relevance: Understanding of the molecular and genetic mechanisms underlying acne and acne scar formation is still in its infancy. However, ongoing research in this area increases our knowledge of disease mechanisms and may contribute to the development of new preventive and treatment strategies. Research confirms the significant role of genetic factors in the development of acne, affecting its occurrence, course and effectiveness of treatment. Particular attention is paid to gene polymorphisms associated with inflammation, androgen metabolism and the immune response, such as CYP17A1 and TNF-α. These data highlight the importance of taking patients' genetic profile into account when diagnosing and choosing therapy, which can improve treatment outcomes and prevent disease relapse. Based on an analysis of literature data, in Uzbekistan there is insufficient understanding of the prognostic significance and role of the NLR and TLR2 genes in the development of acne. This highlights the complexity of the genetic component of acne and indicates the need for additional research to better understand the influence of these and other genetic factors on the pathogenesis of the disease.

Conclusion. An in-depth study of the genetic aspects of acne will not only enrich the scientific understanding of the disease, but will also open up new opportunities for its more effective treatment and prevention.


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ABSTRACT

Relevance: Understanding of the molecular and genetic mechanisms underlying acne and acne scar formation is still

in its infancy. However, ongoing research in this area increases our knowledge of disease mechanisms and may

contribute to the development of new preventive and treatment strategies. Research confirms the significant role of

genetic factors in the development of acne, affecting its occurrence, course and effectiveness of treatment. Particular

attention is paid to gene polymorphisms associated with inflammation, androgen metabolism and the immune

response, such as CYP17A1 and TNF-

α. These data highlight the importance of taking

patients' genetic profile into

account when diagnosing and choosing therapy, which can improve treatment outcomes and prevent disease relapse.

Based on an analysis of literature data, in Uzbekistan there is insufficient understanding of the prognostic significance

and role of the NLR and TLR2 genes in the development of acne. This highlights the complexity of the genetic

component of acne and indicates the need for additional research to better understand the influence of these and

other genetic factors on the pathogenesis of the disease.

Conclusion. An in-depth study of the genetic aspects of acne will not only enrich the scientific understanding of the

disease, but will also open up new opportunities for its more effective treatment and prevention.

KEYWORDS

Acne, molecular genetic mechanisms, kelliod, prognosis.

Research Article

THE ROLE OF GENETIC MUTATIONS IN THE DEVELOPMENT OF ACNE

Submission Date:

July 21, 2024,

Accepted Date:

July 26, 2024,

Published Date:

July 31, 2024

Crossref doi:

https://doi.org/10.37547/ajbspi/Volume04Issue07-05


Khamidova Farida Muinovna

Samarkand State Medical University, Department of Pathological Anatomy with a sectional course, Uzbekistan

Khusinova Firuza Azgar Kizi

Samarkand State Medical University, Department of Pathological Anatomy with a sectional course, Uzbekistan

Journal

Website:

https://theusajournals.
com/index.php/ajbspi

Copyright:

Original

content from this work
may be used under the
terms of the creative
commons

attributes

4.0 licence.


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INTRODUCTION

Acne (L70, L73. 0.) is a chronic inflammatory skin

disease affecting the hair sebaceous follicles and is

caused by a variety of factors including genetics and

androgens. Neonatal acne (acne neonatorum) occurs

in the first four weeks of life, and infantile acne appears

between 3 and 6 months. These conditions may be

associated with high androgen levels in girls and boys

(1,7).

Acne can develop at any age and is often seen in

teenagers and young adults. Symptoms include

comedones, pustules and inflammatory nodules.

Microbial flora, in particular Cutibacterium acnes, plays

a

significant

role

in

pathogenesis,

causing

inflammation and infection.

Treatment for acne includes topical retinoids, benzoyl

peroxide, antibiotics and, in some cases, oral

contraceptives or isotretinoin. It is important to

consider antibiotic resistance and the potential

psychological effects of the disease (2,8).

Historically, acne has been known since ancient times;

its descriptions are found among the ancient Egyptians

and Greeks. Current understanding of the disease

emphasizes the importance of genetic and immune

factors in its development.

Acne is an inflammatory skin disease that can appear at

different points in life and is often associated with

genetic factors (1,2,3,4).

Neonatal, nodular cystic and conglobate acne have a

pronounced genetic predisposition, and acne after

adolescence is associated with a family history of acne

in 50% of cases.

Research has identified genetic markers associated

with acne, including apolipoprotein A1 and various loci

detected through genome-wide analyses. Several

genetic pathways, including PI3K/AKT/mTOR, may

regulate sebum production and inflammation, which

play a key role in the development of acne (5).

Acne severity scoring systems vary and may include

analysis of comedones, papules and pustules.

Immunity research has shown that inflammation in

acne can be caused by a variety of factors, including

skin microbes and innate immune responses.

The following clinical and morphological forms of acne

are distinguished:

L70.0- Common acne (acne vulgaris), L70.1 - Globular

acne, L70.2 Smallpox acne, Necrotic miliary acne, L70.3-

Tropical acne, L70.4-Children's acne, L70.5- Acne

excoriée, Excoriated acne, L70.5-Other acne, L70.9-

Acne, unspecified.

Acne Vulgaris is a globally common chronic

inflammatory disease of the hair and sebaceous

follicles. Although acne is not life-threatening, it can

cause scarring, irritation and serious psychological

problems, including depression. Our review examines

the various causes of acne and methods of treating


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them. Major pathophysiological factors include

excessive sebum production, hyperkeratinization, P.

acnes colonization, and inflammation. In diagnosing

acne, it is important to distinguish between

inflammatory and non-inflammatory forms of lesions.

Problems of antibiotic resistance require the

development of new treatments (12).

Epidemiological studies show that acne affects up to

80% of adolescents and young adults. The age at which

acne most often begins ranges from 14 to 16 years in

girls and 16 to 17 years in boys. However, although acne

affects men and women at approximately the same

frequency, severe forms are more common among

men (17,18,19,20,21,22).

The causes and mechanisms of acne development are

not fully understood, but they are believed to be

associated with a number of factors. These include

hormonal imbalance and hypersecretion of sebum,

changes in its chemical composition, follicular

hyperkeratosis and colonization of the skin by the

bacteria Propionibacterium acnes. An important role in

the development of acne belongs to the immune

response to the antigens of these microorganisms,

which activate neutrophils and phagocytes.

These cells stimulate the complement system through

Toll-like receptors (TLR2), resulting in the synthesis of

pro-inflammatory interleukins such as IL-8, TNF-

α, IL

-

and IL-12. These interleukins activate cyclooxygenase,

which promotes the production of inflammatory

mediators, such as leukotriene B4 from arachidonic

acid. It has been established that leukotriene B4

activates monocytes, eosinophils, T-lymphocytes and

other cells that secrete hydrolytic enzymes that

destroy the wall of the sebaceous gland. This leads to

the release of the contents of the gland into the

surrounding tissues and causes the development of

inflammation at the site of the lesion (23,24).

Immunodeficiencies such as chronic granulomatous

disease (CGD) can aggravate skin conditions such as

folliculitis. CGD is a rare inherited disease caused by a

defect in the enzyme NADPH oxidase, which prevents

white blood cells from effectively killing pathogens.

Such patients may experience recurrent bacterial and

fungal skin infections, including folliculitis and acne.

Diagnosis of CGD involves specific tests for neutrophil

function, and treatment includes infection prevention

and immunomodulators (6,7,8).

Hidradenitis suppurativa, also known as acne inversus

or acne inversus, is a chronic inflammatory skin disease

affecting the hair follicles and is associated with acne.

The disease manifests in intertriginous areas and can

cause significant burden due to pain, itching, malodor,

and emotional distress. Treatment may include

antibiotics, antimicrobials, and surgical interventions

to control symptoms and prevent disease progression

(9,10,11).

A study by Ballanger et al shows a significant influence

of heredity on the development and course of acne. A

family history of acne (A+) is associated with earlier

onset of the disease, often before puberty, and


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increases the recurrence rate after isotretinoin

treatment. In addition, acne patients whose parents

also suffered from acne (M+ and M+F+) are more likely

to experience problems with retention lesions. This

highlights the role of genetic factors as an important

prognostic indicator in clinical practice in the diagnosis

and selection of treatment strategies for acne (15).

Modern research focuses on the genetic aspects of

acne,

indicating

that

genetic

predisposition

significantly influences the occurrence, clinical

presentation, course and effectiveness of acne

treatment. Acne is a polygenic disease, which means

that it does not follow the classical laws of Mednellian

inheritance (25,26).

There are many genetic markers associated with acne,

including polymorphisms in genes such as tumor

necrosis factor alpha, matrix metalloproteinases

(MMPs), tissue inhibitors of metalloproteinases

(TIMPs), interleukin-1, CYP17A1, CYP1A1, and others.

Research by Yaykasli K.O. showed that genotype

frequencies for matrix metalloproteinase-2 (MMP-2)

and TIMP-2 inhibitor polymorphisms were similar in a

Turkish population between acne patients and

controls. However, an imbalance between MMPs and

TIMP-2 may increase susceptibility to acne, as shown

for the TIMP-2 (-418 C/C) genotype, which was twice as

common in patients compared to controls (27).

Most of the genes studied in the context of acne play

key roles in innate immune function, skin lesion

formation, or steroid hormone metabolism. Particular

attention is paid to polymorphisms of genes involved

in the biosynthesis of androgens, including

testosterone, such as cytochrome P450c17α, encoded

by the CYP17A1 gene (28,29,30,31,32).

A study by N. Malikova et al. in the Uzbek population

showed that certain genotypic variants of the CYP17A1

gene are associated with a higher likelihood of

developing acne and its severe course. For example,

the heterozygous A/G genotype was typical for

patients with moderate acne, while the G/G genotype

was more common in patients with severe acne. The

A/A variant was protective and associated with a

reduced risk of acne (33,34).

In China, the CYP17-34T/C polymorphism was

associated with acne, with men with the homozygous

C/C variant and the C allele having a high risk of severe

acne, in contrast to women with mild to moderate

acne, where no such association was observed (35 ,36).

A polymerase chain reaction (PCR) study examined the

effect of CYP17 gene polymorphisms on the

development of acne in Chinese men. The homozygous

C/C variant and the C allele were found to be

significantly more common in men with severe forms

of acne compared to the control group, confirming

statistically significant differences. However, among

women with mild and moderate forms of acne, no

statistically significant differences were found with the

control group, indicating a possible sex difference in

the genetic predisposition to severe forms of acne.

Another study among Indonesian patients showed


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that polymorphisms in the CYP17A1, CYP1A1 and TNF-

α

genes did not correlate with the risk of developing

severe acne. However, it has been found that

polymorphisms in the CYP1A1 gene may contribute to

acne in general. These findings highlight the

complexity of the genetic background of acne and the

need for further research to determine the precise role

of these and other genetic factors in the development

of the disease (37).

Researchers in Germany found that having the GG

genotype of the CYP1A1 gene may increase the risk of

developing acne. This genotype is thought to

contribute to a deficiency of natural retinoids leading

to follicular hyperkeratosis and acne vulgaris. This may

also explain the high effectiveness of retinoids in the

treatment of severe acne in patients with this

polymorphism, since they show a better therapeutic

response than patients without this polymorphism

(38).

Studies have shown that the -308 G/A and -238 G/A

polymorphisms in the TNF gene are significantly more

common in patients with acne vulgaris, especially

those suffering from severe forms of acne, than in

healthy individuals. This is confirmed by the increased

risk of developing acne in the European population

compared to the Asian population. In addition, high

levels of the proinflammatory cytokine IL-8 and a

significant frequency of the IL-8-251T>A polymorphism

are found in Pakistani patients with acne, highlighting

a genetic predisposition to the development and

severity of acne (39,40,41,42).

In addition, analysis of the rs4646421 polymorphism of

the CYP1A1 gene showed that although the activity of

the cytochrome CYP1A1 enzyme is important for the

metabolism of sex hormones and vitamin A, a direct

connection

of

this

polymorphism

with

the

development of acne was not found. However, this

genetic marker has been shown to have significant

prognostic value for predicting severe acne, as patients

with severe acne are more than twice as likely to have

an unfavorable allelic variant as controls (43).

An increase in the level of TSPO in the skin of patients

with acne, along with other enzymes that metabolize

steroids -

3βHSD, CYP11A1, may indicate an

intensification of the synthesis of steroids in the skin

during this period. Pathology, as well as their

contribution to the development of chronic

inflammation in acne (50).

A study from Western Iran found that the PPARγ

Pro12Ala and C161T polymorphisms do not directly

influence the risk of developing acne vulgaris (AV), but

the PPARγ Pro allele is associated with increased

susceptibility to AV in adults over 20 years of age. In

addition, polymorphisms influence the lipid profile of

patients, indicating significantly higher levels of total

cholesterol and triglycerides in carriers of the variant

CG genotype compared to the CC genotype. The study

also found that CT and TT genotypes were associated

with lower serum cholesterol and LDL-C levels. These


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data highlight the importance of genetic factors in the

pathogenesis and clinical presentation of acne, as well

as their possible influence on the lipid profile, which

may have implications for the choice of acne treatment

strategy (16).

A study conducted among Turkish patients with acne

vulgaris found significant differences in the frequency

of the IGF-I (CA)19 genotype between affected and

healthy subjects, supporting its possible influence on

the development of acne (P=0.0002). This genotype

was also found to be associated with acne severity

(P=0.015), suggesting a role in disease progression in

this ethnic group (44,45).

Studies have also shown that tumor necrosis factor

alpha (TNF-

α), a powerful pro

-inflammatory cytokine,

plays a key role in triggering and regulating the

cytokine cascade in inflammation and the immune

response. One of the functional polymorphic loci, G-

308A, has a significant effect on the synthesis and level

of TNF-

α in the div, which emphasizes its potential

importance in the pathogenesis of acne and its clinical

course (46,47).

The authors concluded that elevated levels of

interleukin-8 and its genetic polymorphism IL-8-251T>A

may contribute to the development of acne in the

population. Genetic markers play a key role in the

development and progression of acne. A detailed study

of the genetic and immunogenetic factors associated

with acne may lead to a better understanding of the

molecular and genetic mechanisms of this condition.

This knowledge will help in the development of new

methods to predict the course of the disease and

effective therapeutic approaches to treat acne, taking

into account genetic polymorphisms (48,49).

The formation of scar tissue includes three phases that

follow in a certain time sequence: inflammatory,

proliferative and remodeling phase (51).

The main structural units active in the inflammatory

and proliferative phase are fibroblasts, capillary

endothelium, transform-transient growth factor (TGF)

β1 and β2, platelet

-derived growth factor (PDGF),

insulin-like growth factor (IGF-1) and epidermal growth

factor (EGF). Vascular endothelial growth factor

(VEGF), which is produced by epidermal cells, acts as a

positive regulator of angiogenesis. Tissue inhibitors of

metalloproteinases (TIMPs) are endogenous inhibitors

of matrix metalloproteinases (MMPs). Thus, increased

levels of TIMP are presumably associated with

hypertrophic scar formation. Tumor necrosis factor α

(TNF-

α) is an inflammatory cytokine produced by

monocytes and macrophages during the inflammatory

phase. This cytokine is known to cause collagen

degradation and help minimize excessive scarring (52).

During the remodeling phase, excess extracellular

matrix is degraded and type III collagen, the immature

form of collagen, is replaced by mature type I collagen.

TGF-

β3 is believed to play a leading role in this process.

Also, members of the MMP family have a significant

effect on the degradation and remodeling of the ECM

and mediate the degradation of collagen types I and III,


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reducing inflammation and neutralizing the effects of

chemokines (53).

Decorin is a proteoglycan component of cutaneous

connective tissue that binds to type I collagen fibrils

and influences TGF-

β. By binding and neutralizing TGF

-

β, decorin reduces the stimulating effect

of TGF-

β on

the

synthesis

of

collagen,

fibronectin

and

glycosaminoglycan. Decorin levels are reduced in

keloids and hypertrophic scars, and its antifibrotic

properties have attracted attention as a possible

therapeutic agent (54,55).

The role of periosteum (extracellular matrix protein),

the level of which is sharply increased in hypertrophic

scars and keloids compared to normal tissues, is also

actively discussed (56,57).

Proinflammatory factors such as interleukins IL-

1α, IL

-

1β, IL

-6, and tumor necrosis factor alpha are

upregulated in keloid tissues, suggesting that patients

with

keloids

have

increased

expression

of

proinflammatory genes in the skin. This may contribute

to chronic inflammation, which, in turn, can cause

invasive growth of keloids, although according to the

results of a study that was conducted in Turkey (90

people in the study group and 30 in the control group),

polymorphic variants

Ants TNF-

α (

-308 G/A) and IL-

1β (

-511 C/T) were not

associated with acne susceptibility, acne scarring, or

acne severity (58).

Increased expression of pro-inflammatory factors

means that keloids and hypertrophic scars are a

consequence of inflammatory processes in the

reticular layer of the dermis. Various external and

internal stimuli (local, systemic and genetic) after injury

can contribute to inflammation. The nature of these

irritants most likely determines the characteristics,

number, and development of keloids and hypertrophic

scars. In England, when studying a small group of

patients with keloid scars (including acne), they found

that the presence of HLADRB5 and HLA-DRB1*15 was

associated with keloid disease (59).

A meta-analysis of the association of the Arg72Pro

polymorphism of the P53 gene with keloid scars in the

Chinese population included the results of 6 studies,

which included 359 patients with keloid scars and 493

people. as a control. It was determined that the Pro

allele of the Arg72Pro polymorphism of the P53 gene is

a risk factor for the development of keloids in the

Chinese population compared to the Arg allele (OR =

2.29, 95% CI = 1.45

3.60) (60).

A study was conducted in China in which the first stage

analyzed 1056 patients with acne and 1056 controls

using high-density chips. At the second stage of the

study, in an independent cohort (1860 patients and

3660 people in the control group), 101 single

nucleotide polymorphisms were tested, of which 3

showed an association: rs747650 of the DDB2 gene and

rs1060573 (11p11.2), rs7531806 of the SELL gene

(1q24.2), which are involved in androgen metabolism,

inflammation, and scar formation in severe acne (61).


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Keloids develop in fibroproliferative disorders against

the background of chronic inflammatory processes in

the skin. A genome-wide association study showed an

association with rs8032158 in the NEDD4 gene. This

gene is expressed by neural progenitor cells and has six

different transcripts. Carriage of the rs8032158 risk

allele C in patients with keloids is associated with

selectively higher expression of type 3 transcript (TV3

NEDD4) and activation of the NF-

κB pathway. The

analysis revealed that NEDD4 TV3 is involved in NF-

κB

activation through its association with the adapter

protein RIP. These results suggest that NEDD4 TV3 is a

potential diagnostic marker and therapeutic target for

chronic skin diseases, including keloid (62).

A genome-wide association study of 478 African

Americans (122 cases, 356 controls) was conducted in

the USA in 2014. An association was found with the

q21.2-22.3 locus on chromosome 15, which includes the

NEDD4 gene. This gene has previously been shown to

be associated with keloid scars in Japanese and

Chinese populations. But in African Americans, a more

significant association was found with the MYO1E

gene. In addition, an association was established with

the q13.5 locus on chromosome 11 (MYO7A gene,

rs35641839, OR = 4.71, 95% CI 2.38

9.32, p = 8.34 × 10

6). The authors suggest that the identification of

polymorphisms associated with the formation of

keloid scars in two myosin genes indicates that the

altered cytoskeleton contributes to enhanced

migratory and invasive properties of keloid fibroblasts

(63).

Long non-coding RNAs (lncRNAs) are believed to play

a significant role in human diseases. Studies have

shown that overexpression of long non-coding RNA

AC067945.2 did not affect cell proliferation in

hypertrophied scar tissue, but promoted early

apoptosis in normal skin fibroblasts. Except Moreover,

overexpression

of

AC067945.2

inhibited

the

expression of COL1A1, COL1A2, COL3A1 and α

-SMA

proteins. In turn, TGF-

β1 can inhibit t

he expression of

AC067945.2. In the group with overexpression of

AC067945.2, 138 mRNA expression differed from the

control group, of which it was increased in 14 and

decreased in 124. Overexpression of AC067945.2

correlated with developmental processes, binding,

extracellular region and the VEGF and Wnt signaling

pathways. The study revealed the functions of the

novel lncRNA AC067945.2, which may help understand

the mechanisms regulated by AC067945.2 in the

pathogenesis of hypertrophic scars (64).

Overexpression of long non-coding RNA ncRNA8975-1

was found in hypertrophic scars and skin fibroblasts.

Overexpression of lncRNA8975-1 prevents cell

proliferation and reduces the expression of COL1A2,

COL1A1, COL3A1 and α

-SMA in hypertrophic scar

fibroblasts, whereas knockdown of lncRNA8975-1 has

the opposite effect. Further studies of the mechanisms

by which lncRNA8975-1 expression is regulated may


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lead to a better understanding of the pathogenesis of

hypertrophic scars (65).

The study of the genetic aspects of the problem of scar

formation after acne, as well as the determination of

molecular genetic markers of this condition, is almost

at the very initial stage. It is likely that this gap will be

filled in the coming years, which will provide impetus

for the development of new effective means of

preventing and treating this disease (71). regulators of

androgen receptors - 25-OH-VD, cytochrome p 450 (17-

alpha hydroxylase), insulin-like growth factor are

important in the pathogenesis of rapid non-genomic

molecular cellular reactions of peripheral androgen

metabolism and, in the future, may determine new

algorithms for the diagnosis and treatment of acne

diseases (66).

An important aspect of acne pathogenesis is the

participation of keratinocytes in the inflammatory

response. C. acnes activates Toll-like receptor (TLR)-2

and TLR-4 on keratinocytes, leading to activation of

signaling cascades including the NF-

κB pathway and

the MAPK pathway. Subsequently, keratinocytes

produce IL-1, IL-8, IL-6, granulocyte-macrophage

colony-stimulating factor (GM-CSF), TNF-

α, matrix

metalloproteinases (MMPs) and human β

-defensin-2

(hBD-2). In addition to TLR-2 and TLR-4, the CD36

receptor expressed on keratinocytes is also involved in

the recognition of C. acnes.

Once C. acnes CD36 is detected, keratinocytes begin to

synthesize reactive oxygen species (ROS), especially

superoxide anion, generated from the cytosolic

enzymes NAD(P)H oxidases. These ROS provide an

antibacterial effect and trigger an inflammatory

response. Analysis of identified variants of the

nucleotide sequence of keratinocyte proliferation and

differentiation genes showed that severe acne is likely

associated with polymorphic loci AP3B1, FERMT1,

FERMT3, GBA, SUFU (67).

Data on the role of family history and possible

inheritance of acne have been confirmed in a number

of studies on familial cases of this dermatosis, its more

frequent development in monozygotic twins, but the

significance of genetic associations is not fully known.

Thus, He L. et al. (2014) revealed a connection between

acne and TP63, which ensures the regulation and

differentiation of epithelial stem cells (68).

In another study, the authors suggested an

association of acne development with LGR6, which is a

mediator of the WNT signaling pathway and ensures

the functioning and differentiation of sebaceous gland

stem cells. Additional associated genes were LAMC2,

encoding a major component of the basement

membrane, and SPECC1L, encoding a cross-linking

cytoskeletal protein that plays an important role in cell

adhesion and migration (69).

It has been shown that the presence of the disease in

first-degree relatives may be a risk factor for the

development of dermatosis. A case of identification

and determination of the significance of polymorphism

of the NCF1, CD3E, ORAI1, IGHM, TAZ genes in patients


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with severe forms of the disease with a family history

is presented. The conducted studies revealed identical

allelic variants in five genes: NCF1, CD3E, ORAI1, IGHM,

TAZ in two closely related patients (father and son)

with severe acne. Polymorphisms of the studied genes

probably influence the development of an imbalance in

the oxidase system, the functioning of mitochondria,

reduced proliferation of T cells, as well as the

formation of an imbalance in the secretion of

immunoglobulins. The data obtained may be factors in

the torpid course of severe dermatosis, which

determines the need for further research (70). Based

on the analysis of literature data, it can be noted that

in our Republic of Uzbekistan, the prognostic values,

features and role of the NLR and TLR2 genes, and their

relationship in the development of acne have not been

fully studied.

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НОВЫЕ

ПАТОГЕНЕТИЧЕСКИЕ

ФАКТОРЫ

АНДРОГЕНЗАВИСИМЫХ

ДЕРМАТОПАТИЙ.

д.м.н. Азимова Ф. В., Ходжаева М. Б.

International Academy Journal Web of Scholar

7(37), July 2019.

67.

ПАТОГЕНЕТИЧЕСКАЯ

РОЛЬ

АЛЛЕЛЕЙ

ПОЛИМОРФНЫХ

ВАРИАНТОВ

ГЕНОВ

ПРОЛИФЕРАЦИИ И ДИФФЕРЕНЦИРОВКИ

КЕРАТИНОЦИТОВ ПРИ ТЯЖЕЛОЙ СТЕПЕНИ

АКНЕ.Демина О.М., Международный научно

-

исследовательский журнал

1 (127)

Январь

.

68.

He L. Two new susceptibility loci 1q24.2 and

11p11.2 confer risk to severe acne / L. He, W.J.

Wu, J.K. Yang et al. //Nat Commun.

2014.

5.

p. 2870.

69.

Petridis

C.

Genome-wide

meta-analysis

implicates

mediators

of

hair

follicle

development and morphogenesis in risk for


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American Journal Of Biomedical Science & Pharmaceutical Innovation
(ISSN

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VOLUME

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Publisher:

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severe acne / C. Petridis, A.A. Navarini, N. Dand

et al. // Nat Commun.

2018.

9.

p.5075.

70.

РОЛЬ ГЕНЕТИЧЕСКИХ ФАКТОРОВ ПРИ

СЕМЕЙНОМ СЛУЧАЕ АКНЕ О. М. Демина, А.

Г. Румянцев, Н. Н. Потекаев.ВЕСТНИК РГМУ

3, 2022 стр.36

-39.

71.

Nemchaninova O.B. et al. / Journal of Siberian

Medical Sciences 2 (2020) 98

110

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НОВЫЕ ПАТОГЕНЕТИЧЕСКИЕ ФАКТОРЫ АНДРОГЕНЗАВИСИМЫХ ДЕРМАТОПАТИЙ. д.м.н. Азимова Ф. В., Ходжаева М. Б. International Academy Journal Web of Scholar 7(37), July 2019.

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